Orobanche californica subsp. grayana

(Beck) Heckard

Madroño 22: 54. 1973.

Endemic
Basionym: Orobanche grayana Beck Biblioth. Bot. 19: 79. 1890
Synonyms: Aphyllon californicum subsp. grayanum (Beck) A. C. Schneider Myzorrhiza grayana (Beck) Rydberg
Treatment appears in FNA Volume 17. Treatment on page 480. Mentioned on page 478.

Plants usually branched proximally, 4–10 cm, portion proximal to inflorescence 1–3 (–4) cm, slender. Inflorescences corymbs, sometimes subcorymbose racemes, 3–6 cm; bracts pallid to pinkish tinged, drying brown. Pedicels 5–20 mm. Calyces 11–15 mm, lobes pallid, sometimes purplish, linear-subulate, (7–) 9–13 (–16) mm. Corollas pallid to pinkish or pale lavender, often with lavender veins, (25–) 28–33 mm; tube slender, abruptly widening toward throat; throat 8–10 mm wide at base of lobes; lips 10–12 mm, abaxial lobes lanceolate to lanceolate-subulate, 2–3 mm wide, apex acute, adaxial lobes oblong-ovate, apex narrowly rounded, shallowly retuse, or erosulate. 2n = 48.


Phenology: Flowering (Jun–)Aug–Sep.
Habitat: Moist meadows and stream margins.
Elevation: (50–)300–2100 m.

Discussion

Subspecies grayana is distributed in the Cascade-Sierra Nevada ranges, Coast Ranges (central California and southern Oregon), and mountains of the Columbia Plateau in Oregon, from Klickitat County, Washington, south to Tuolumne County, California. It is rare throughout the range, possibly locally extirpated in portions of the range in Oregon.

The hosts are primarily species of Aster and Erigeron and occasionally Grindelia. Other reported non-Asteraceae hosts are unlikely.

Selected References

None.

Lower Taxa

None.
lanceolate +  and lanceolate-subulate +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
oblong-ovate +
glabrous +  and villous +
glandular-pubescent +
retuse +, rounded +  and acute +
L. Turner Collins +, Alison E. L. Colwell +  and George Yatskievych +
(Beck) Heckard +
enlarged +
Orobanche grayana +
oblong;lanceolate;oblong;lanceolate;deltate;triangular;ovate +
0.4 cm4 mm <br />0.004 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
not leathery +, not +  and fleshy +
brown;pallid;pinkish tinged +
appressed +  and spreading +
less lanceolate +  and oblanceolate +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
2 cm20 mm <br />0.02 m <br /> (2.7 cm27 mm <br />0.027 m <br />) +
pallid +  and dark purple pink yellow or white +
divided +
1.1 cm11 mm <br />0.011 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
ovoid;cylindric-ovoid +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
2.5 cm25 mm <br />0.025 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
pale lavender +, pallid +  and pinkish +
bent +  and tubular +
constricted +
2.8 cm28 mm <br />0.028 m <br /> (3.3 cm33 mm <br />0.033 m <br />) +
spikelike +
Calif. +, Oreg. +  and Wash. +
(50–)300–2100 m. +
pubescent +  and glabrous +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Moist meadows and stream margins. +
3 cm30 mm <br />0.03 m <br /> (4 cm40 mm <br />0.04 m <br />) +
axillary +  and terminal +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
branched +  and simple +
pallid cream +  and nearly white +
deciduous +
cream +  and pinkish or purplish tinged or pink +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
1.3 cm13 mm <br />0.013 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
purplish +  and pallid +
loosely ascending +  and recurved +
glandular-pubescent +
linear-subulate +
erosulate +  and entire +
lobed +, toothed +  and entire +
tenuinucellate +  and unitegmic +
campylotropous-like +
spikelike +
2 cm20 mm <br />0.02 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Flowering (Jun–)Aug–Sep. +
subcorymbose +
spikelike +
3 cm30 mm <br />0.03 m <br /> (6 cm60 mm <br />0.06 m <br />) +
branched +  and unbranched +
inconspicuous +
coralloid +
stout +  and slender +
black;tan;dark-brown +
500 +  and 2000 +
prismatic +, ovoid +  and oblong-ellipsoid +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
5 +  and 4 +
subequal +
yellow +  and white +
aerial +  and subterranean +
puberulent +  and glabrous +
capitate +  and crateriform-peltate +
clavate +  and 2-4-lobed +
Aphyllon californicum subsp. grayanum +  and Myzorrhiza grayana +
Orobanche californica subsp. grayana +
Orobanche californica +
subspecies +
8mm;10mm +
cream;pinkish or purplish tinged or pink +
glandular-puberulent +
slender +
widening +
35 cm350 mm <br />0.35 m <br /> (40 cm400 mm <br />0.4 m <br />) +
perennial +  and annual +
holoparasitic +  and hemiparasitic +
stout +  and slender +