Parrya nudicaulis

(Linnaeus) Regel

Bull. Soc. Imp. Naturalistes Moscou 34: 176. 1861.

Basionym: Cardamine nudicaulis Linnaeus Sp. Pl. 2: 654. 1753
Synonyms: Achoriphragma nudicaule (Linnaeus) Soják Arabis nudicaulis (Linnaeus) de Candolle Cardamine articulata Pursh Matthiola nudicaulis (Linnaeus) Trautvetter Neuroloma nudicaule (Linnaeus) Andrzejowski ex de Candolle Parrya macrocarpa R. Brown Parrya nudicaulis var. grandiflora Hultén Parrya nudicaulis subsp. interior Hultén Parrya nudicaulis var. interior (Hultén) B. Boivin Parrya nudicaulis subsp. septentrionalis Hultén
Treatment appears in FNA Volume 7. Treatment on page 512. Mentioned on page 513, 514.

Plants often not cespitose, caudex branched; glandular throughout or eglandular. Stems (0.4–) 0.7–2.7 (–3.5) dm. Leaves: petiole (0.7–) 1.5–7 (–10) cm, to 5 mm wide at base (glandular or not); blade narrowly spatulate or oblanceolate to lanceolate or oblong, (1–) 1.7–7 cm × (6–) 10–23 (–28) mm, base cuneate or attenuate, margins entire or minutely to coarsely dentate, or sometimes incised, apex acute. Racemes 3–20-flowered. Fruiting pedicels (proximalmost) (10–) 15–40 (–60) mm (glandular or not). Flowers: sepals oblong, 5–8 × 1.5–3 mm (glandular or not); petals lavender to white, purple, (14–) 16–20 (–22) × 7–10 (–12) mm, claw 6–10 mm, apex emarginate; median filaments 6–10 mm; anthers 1.5–2.5 mm. Fruits narrowly oblong to linear-lanceolate, (2–) 3–4 (–4.7) cm × (3.5–) 5–7 mm; valves glandular or eglandular; ovules 12–16 per ovary; style (0.5–) 1–2.5 (–3.5) mm. Seeds 3.5–6 × 3–5 mm; wing 0.7–1.5 mm wide. 2n = 14, 28.


Phenology: Flowering early Jun-early Aug.
Habitat: Tundra, alpine stream valleys, flats and flood banks, limestone or schist mountain slopes and tops, grassy summits, disturbed gravel, moist open areas, meadows, sandy shores, mossy carpets, hillsides, alpine stony slopes, stable sand ridges, turfy snow flushes
Elevation: 0-1800 m

Distribution

V7 806-distribution-map.gif

B.C., N.W.T., Yukon, Alaska, e Asia (Russian Far East)

Discussion

Parrya nudicaulis is the most variable species in the genus, especially in leaf shape, size, and margin, as well as in the presence versus absence of the extrafloral glands. Much of the confusion about its limits resulted from different emphases on various characters. For example, E. Hultén (1971), who recognized six subspecies, expanded the range of P. nudicaulis to extend from the Canadian arctic and Alaska into the Russian Far East, Siberia, Central Asia, China, and the Himalayas. Three of his four North American subspecies, sometimes growing together, are rather poorly defined morphologically and appear to have been based primarily on the degree of development of leaf teeth. By contrast, R. C. Rollins (1993) recognized a single polymorphic species that included the Utah-Wyoming endemic P. rydbergii. Within a given population of P. nudicaulis, one finds both glandular and eglandular plants with leaf margins entire or variously dentate. In my opinion, these variables alone are unreliable, and flower size, in combination with other characters, can give a better indication of taxon identity.

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Parrya nudicaulis"
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
Ihsan A. Al-Shehbaz +
(Linnaeus) Regel +
attenuate +  and cuneate +
Cardamine nudicaulis +
appendaged +  and unappendaged +
1 cm10 mm <br />0.01 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
2.3 cm23 mm <br />0.023 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
not auriculate +
1.7 cm17 mm <br />0.017 m <br /> (7 cm70 mm <br />0.07 m <br />) +
dentate +, subentire +  and entire +
oblanceolate;lanceolate or oblong +
much smaller +  and reduced +
1 cm10 mm <br />0.01 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
urceolate +, campanulate +  and tubular +
eglandular +  and branched +
well-developed +
distinct +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
differentiated +
emarginate +  and entire +
B.C. +, N.W.T. +, Yukon +, Alaska +  and e Asia (Russian Far East) +
0-1800 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
not basally +  and dilated +
actinomorphic +
, +, latiseptate +, stipitate +  and sessile +
4 cm40 mm <br />0.04 m <br /> (4.7 cm47 mm <br />0.047 m <br />) +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
indehiscent +  and dehiscent +
3 cm30 mm <br />0.03 m <br /> (4 cm40 mm <br />0.04 m <br />) +
narrowly oblong +  and linear-lanceolate +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
Tundra, alpine stream valleys, flats and fTundra, alpine stream valleys, flats and flood banks, limestone or schist mountain slopes and tops, grassy summits, disturbed gravel, moist open areas, meadows, sandy shores, mossy carpets, hillsides, alpine stony slopes, stable sand ridges, turfy snow flusheses, stable sand ridges, turfy snow flushes +
connate +  and distinct +
decurrent +
dentate +  and entire +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
semiannular +  and annular +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
2 cm20 mm <br />0.02 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
lavender +  and white +
16mm;20mm +
rudimentary +
obovate +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
7 cm70 mm <br />0.07 m <br /> (10 cm100 mm <br />0.1 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (7 cm70 mm <br />0.07 m <br />) +
0 cm0 mm <br />0 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Flowering early Jun-early Aug. +
trinucleate +  and 3(-11)-colpate +
Bull. Soc. Imp. Naturalistes Moscou +
3-20-flowered +
4 cm40 mm <br />0.04 m <br /> (6 cm60 mm <br />0.06 m <br />) +
divaricate-ascending +  and ascending +
1.5 cm15 mm <br />0.015 m <br /> (4 cm40 mm <br />0.04 m <br />) +
flattened +
mucilaginous +
aseriate +, biseriate +  and uniseriate +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
flattened +, suborbicular +  and broadly ovate +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
persistent +  and caducous +
distinct +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
procumbent +  and decumbent +
2-lobed +, cylindric +  and conical +
subsessile +, sessile +  and petiolate +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
persistent +
distinct +
0.1 cm1 mm <br />0.001 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
Achoriphragma nudicaule +, Arabis nudicaulis +, Cardamine articulata +, Matthiola nudicaulis +, Neuroloma nudicaule +, Parrya macrocarpa +, Parrya nudicaulis var. grandiflora +, Parrya nudicaulis subsp. interior +, Parrya nudicaulis var. interior +  and Parrya nudicaulis subsp. septentrionalis +
Parrya nudicaulis +
species +
scale-like +  and forked +
eglandular +  and glandular +
pubescent +  and glabrous +
anastomosing +
eglandular +, glandular +  and scapose +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
plant +  and cespitose +
aquatics +  and terrestrial +
glabrous +  and pubescent +
0.7mm;1.5mm +