Penstemon arkansanus

Pennell

Proc. Acad. Nat. Sci. Philadelphia 73: 493. 1922.

Common names: Arkansas beardtongue
Endemic
Synonyms: Penstemon multicaulis Pennell P. pallidus subsp. arkansanus (Pennell) R. W. Bennett P. wherryi Pennell
Treatment appears in FNA Volume 17. Treatment on page 193. Mentioned on page 185, 186, 215, 224.

Herbs. Stems ascending to erect, (18–) 25–45 (–61) cm, retrorsely hairy, sometimes also ± glandular-pubescent distally, not glaucous. Leaves basal and cauline, basal often withering by anthesis, not leathery, glabrous or sparsely retrorsely hairy, especially along midvein and margins, abaxially, glabrous or sparsely puberulent adaxially; basal and proximal cauline 20–95 × 5–28 mm, blade spatulate or obovate to ovate, base tapered, margins subentire to ± serrate or dentate, apex rounded to obtuse or acute; cauline (6–) 8–12 pairs, short-petiolate or sessile, 22–80 × 4–20 mm, blade lanceolate, base tapered to clasping, margins ± serrate or dentate, apex acute to acuminate. Thyrses interrupted, conic, 6–24 cm, axis retrorsely hairy and ± glandular-pubescent, verticillasters 4–8, cymes 3–13-flowered, 2 per node; proximal bracts lanceolate, 6–40 × 2–12 mm, margins entire or ± dentate; peduncles and pedicels ascending, retrorsely hairy and ± glandular-pubescent. Flowers: calyx lobes ovate, 2.8–4.5 × 1.1–2 mm, sparsely glandular-pubescent; corolla white, with reddish purple nectar guides, tubular to tubular-funnelform, 15–19 mm, glandular-pubescent externally, moderately whitish lanate internally abaxially, tube 4–6 mm, throat slightly inflated, 4–7 mm diam., 2-ridged abaxially; stamens included, pollen-sacs opposite, navicular, 0.8–1 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate; staminode 10–12 mm, exserted, 0.2–0.4 mm diam., tip straight to slightly recurved, distal 5–7 mm ± villous, hairs yellow or golden yellow, to 1.2 mm; style 9–11 mm. Capsules 4–7 × 4–5 mm, glabrous. 2n = 16.


Phenology: Flowering Apr–Jun.
Habitat: Rocky oak-hickory woodlands, glades, rocky stream bottoms.
Elevation: 60–300 m.

Discussion

Penstemon arkansanus is centered in the Interior Highlands. The species is most similar to P. pallidus, from which it differs by its narrower, less hairy, and more numerous cauline leaves. Some specimens from the narrow zone of sympatry in north-central Arkansas and south-central Missouri exhibit varying degrees of morphologic intermediacy. Penstemon arkansanus has been reported from Illinois (R. H. Mohlenbrock 1986) and Texas (S. D. Jones et al. 1997); those reports seem unlikely.

Selected References

None.

Lower Taxa

None.

"/4+timescorollathroat" is not declared as a valid unit of measurement for this property.

... more about "Penstemon arkansanus"
acute +  and acuminate +
Craig C. Freeman +
Pennell +
glandular-pubescent +  and hairy +
alternate +, subopposite +, whorled +  and opposite +
glaucescent +
alternate +, opposite +, whorled +, helical +  and subopposite +
cauline +, proximal +  and basal +
rarely +, not glaucous +, glabrate +, pubescent +, hairy +, glandular-lanate +, glandular-pubescent +, puberulent +, glandular +  and glabrous +
puberulent +, glabrous +  and hairy +
not leathery +
not +  and leathery +
tapered;tapered +
lanceolate;obovate;ovate +
not +  and leathery +
2.8mm;4.5mm +
glandular-pubescent +
short-tubular +  and short-campanulate +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
sessile +  and short-petiolate +
2.2 cm22 mm <br />0.022 m <br /> (8 cm80 mm <br />0.08 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (2 cm20 mm <br />0.02 m <br />) +
petiolate +  and sessile +
Arkansas beardtongue +
personate +, not personate +, bilabiate +  and symmetric +
whitish +  and white +
lanate +  and glandular-pubescent +
tubular +  and tubular-funnelform +
ampliate +
1.5 cm15 mm <br />0.015 m <br /> (1.9 cm19 mm <br />0.019 m <br />) +
3-13-flowered +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
scale-like +  and not needlelike +
Ark. +, Mo. +  and Okla. +
drupe-like +
60–300 m. +
curved +  and straight +
1 +  and 4 +
Rocky oak-hickory woodlands, glades, rocky stream bottoms. +
golden yellow +  and yellow +
0 cm0 mm <br />0 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
membranous +
terminal +  and axillary +
persistent +  and deciduous +
cauline +  and basal +
orbiculate;linear;orbiculate;linear;truncate;oblanceolate;obovate;oblong;elliptic;round;lanceolate;ovate +
dentate +, entire +  and serrate +
glutinous +
glandular-ciliolate +, hairy +, puberulent +, glabrescent +, glabrous +  and glandular-pubescent +
subentire to more +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
glandular-pubescent +  and hairy +
glandular-pubescent +  and hairy +
5 +  and 4 +
Flowering Apr–Jun. +
navicular-sigmoid +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
1/5 +  and 2/3 +
0.6 cm6 mm <br />0.006 m <br /> (4 cm40 mm <br />0.04 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
Proc. Acad. Nat. Sci. Philadelphia +
minute +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
black +, gray +, brown +  and tan +
angled +  and slightly rounded +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
1 cm10 mm <br />0.01 m <br /> (?) +  and 1.2 cm12 mm <br />0.012 m <br /> (?) +
subalternate +, alternate +, whorled +  and opposite +
45 cm450 mm <br />0.45 m <br /> (61 cm610 mm <br />0.61 m <br />) +
glutinous +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
ascending;erect +
not glaucous +, glandular-pubescent +  and hairy +
25 cm250 mm <br />0.25 m <br /> (45 cm450 mm <br />0.45 m <br />) +
0.9 cm9 mm <br />0.009 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
papillate +
denticulate +
Penstemon multicaulis +, P. pallidus subsp. arkansanus +  and P. wherryi +
Penstemon arkansanus +
Penstemon sect. Penstemon +
species +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
2-ridged +  and inflated +
not constricted +
6 cm60 mm <br />0.06 m <br /> (24 cm240 mm <br />0.24 m <br />) +
straight +
gibbous +  and not spurred +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
cauline +  and proximal +