Penstemon bicolor

(Brandegee) Clokey & D. D. Keck

Bull. S. Calif. Acad. Sci. 38: 12. 1939.

Common names: Two-color beardtongue
Endemic
Basionym: Penstemon palmeri var. bicolor Brandegee Univ. Calif. Publ. Bot. 6: 360. 1916
Synonyms: P. bicolor subsp. roseus Clokey & D. D. Keck P. pseudospectabilis subsp. bicolor (Brandegee) D. D. Keck
Treatment appears in FNA Volume 17. Treatment on page 247. Mentioned on page 248.

Herbs. Stems ascending to erect, 60–150 cm, glaucous. Leaves: basal and proximal cauline 37–110 × 10–50 mm, blade obovate, base tapered, margins coarsely serrate to coarsely dentate, apex obtuse to acute; cauline 4–8 pairs, sessile, 40–110 × 15–55 mm, blade ovate, base connate-perfoliate, margins coarsely serrate to coarsely dentate, apex acute to acuminate. Thyrses interrupted, secund, 16–90 cm, axis glandular-pubescent, verticillasters 9–23, cymes 1–4-flowered; proximal bracts ovate to lanceolate, 8–25 × 3–28 mm; peduncles and pedicels ascending to erect, glandular-pubescent. Flowers: calyx lobes ovate, 4–6 × 2.2–3 mm, glandular-pubescent; corolla pink to rose-pink, purple, or light yellow, with or without reddish or reddish purple nectar guides, strongly bilabiate, ventricose, 18–27 mm, glandular-pubescent externally, glandular-pubescent internally, and sparsely white or yellowish lanate internally abaxially, tube 7–10 mm, length 1.7–2 times calyx lobes, throat abruptly inflated, constricted at orifice, 6–9 mm diam., rounded abaxially; stamens: longer pair reaching orifice, filaments of shorter pair glandular-puberulent proximally, pollen-sacs explanate, 1.4–1.8 mm, sutures smooth; staminode 14–16 mm, included or exserted, 0.3–0.4 mm diam., tip recurved, distal 4–5 mm lanate, hairs yellow, to 2.8 mm, proximal 3–4 mm glandular-puberulent; style 14–16 mm, glabrous. Capsules 10–13 × 5–7 mm, glabrous or sparsely glandular-pubescent distally.


Phenology: Flowering Mar–Jun.
Habitat: Gravelly soils, arroyos, roadsides, talus slopes, desert scrub, juniper woodlands.
Elevation: 500–1700 m.

Discussion

Penstemon bicolor is known from the Mojave Desert in the Black Mountains of northwestern Mohave County, Arizona, the Castle, Clark, and New York mountains of eastern San Bernardino County, California, and the desert mountain ranges of southern and western Clark County, Nevada. Two subspecies differing in corolla color have been recognized, with yellow-corolla forms restricted to Clark County, Nevada. Subspecies are not recognized here, following J. D. Morefield (2006).

Habitat destruction and genetic swamping caused by hybridization between Penstemon bicolor and P. palmeri threaten many populations of P. bicolor in the vicinity of Las Vegas, Nevada (G. Glenne 2003).

Lower Taxa

None.

"/4+timescorollathroat" is not declared as a valid unit of measurement for this property."-2timescalyxlobes" is not declared as a valid unit of measurement for this property.

... more about "Penstemon bicolor"
acute +  and acuminate +
Craig C. Freeman +
(Brandegee) Clokey & D. D. Keck +
glandular-pubescent +
alternate +, opposite +, whorled +, helical +  and subopposite +
connate-perfoliate +
tapered +
Penstemon palmeri var. bicolor +
ovate;obovate +
not +  and leathery +
glandular-pubescent +
short-tubular +  and short-campanulate +
1 cm10 mm <br />0.01 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
glandular-pubescent +  and glabrous +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
4 cm40 mm <br />0.04 m <br /> (11 cm110 mm <br />0.11 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
Two-color beardtongue +
not personate +, bilabiate +  and strongly bilabiate +
pink +  and rose-pink purple or light yellow +
hairy +, glandular-pubescent +  and glabrous +
cylindric-funnelform +, tubular-funnelform +  and ventricose +
ventricose-ampliate +
1-4-flowered +
scale-like +  and not needlelike +
Ariz. +, Calif. +  and Nev. +
drupe-like +
500–1700 m. +
curved +  and straight +
glandular-puberulent +
1 +  and 4 +
Gravelly soils, arroyos, roadsides, talus slopes, desert scrub, juniper woodlands. +
0 cm0 mm <br />0 m <br /> (0.28 cm2.8 mm <br />0.0028 m <br />) +
membranous +
terminal +  and axillary +
persistent +  and deciduous +
3.7 cm37 mm <br />0.037 m <br /> (11 cm110 mm <br />0.11 m <br />) +
cauline +, proximal +  and basal +
glaucous +  and glabrous +
leathery +
1 cm10 mm <br />0.01 m <br /> (5 cm50 mm <br />0.05 m <br />) +
orbiculate;linear;orbiculate;linear;truncate;oblanceolate;obovate;oblong;elliptic;round;lanceolate;ovate +
erose +  and entire +
dentate +  and serrate +
glandular-pubescent +  and glabrous +
denticulate +, toothed +  and entire +
serrate to coarsely +
reddish purple +  and reddish +
1.7-2 times calyx lobes +
glandular-pubescent +
rounded +, inflated +  and ventricose +
constricted +
0.6 cm6 mm <br />0.006 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
ascending +  and erect +
glandular-pubescent +
ascending;erect +
glandular-pubescent +
5 +  and 4 +
Flowering Mar–Jun. +
divergent +  and opposite +
navicular-sigmoid +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
denticulate +  and navicular +
0.8 cm8 mm <br />0.008 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
ovate +  and lanceolate +
0.3 cm3 mm <br />0.003 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
glandular-puberulent +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
Bull. S. Calif. Acad. Sci. +
glenne2003a +  and morefield2006a +
minute +
brown +  and dark-brown or black +
rounded +  and angled +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
exserted +  and included +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
exserted +  and included +
1.4 cm14 mm <br />0.014 m <br /> (?) +  and 1.6 cm16 mm <br />0.016 m <br /> (?) +
subalternate +, alternate +, whorled +  and opposite +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
ascending;erect +
60 cm600 mm <br />0.6 m <br /> (150 cm1,500 mm <br />1.5 m <br />) +
1.4 cm14 mm <br />0.014 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
papillate +
denticulate +
P. bicolor subsp. roseus +  and P. pseudospectabilis subsp. bicolor +
Penstemon bicolor +
Penstemon sect. Spectabiles +
species +
yellowish +  and white +
1.7-2 times calyx lobes +
rounded +  and inflated +
constricted +
0.6 cm6 mm <br />0.006 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
secund +  and interrupted +
cylindric +
16 cm160 mm <br />0.16 m <br /> (90 cm900 mm <br />0.9 m <br />) +
straight +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0 cm0 mm <br />0 m <br /> (0.01 cm0.1 mm <br />1.0e-4 m <br />) +
gibbous +  and not spurred +
yellowish +  and white +
rounded +  and inflated +
constricted +
0.6 cm6 mm <br />0.006 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +