Penstemon californicus

(Munz & I. M. Johnston) D. D. Keck

Bull. Torrey Bot. Club 64: 378. 1937.

Common names: California beardtongue
Basionym: Penstemon linarioides var. californicus Munz & I. M. Johnston Bull. S. Calif. Acad. Sci. 23: 31. 1924
Treatment appears in FNA Volume 17. Treatment on page 101. Mentioned on page 99.

Stems ascending to erect, (5–) 8–30 cm, retrorsely hairy, hairs white, scalelike. Leaves ± leathery or not, densely retrorsely hairy, hairs appressed, white, scalelike; cauline 10–30 pairs, petiolate, 8–16 × 1.5–2.5 mm, blade oblanceolate, base tapered, apex mucronate. Thyrses interrupted or continuous, secund, (1–) 2–9 cm, axis retrorsely hairy with appressed, white, scalelike hairs, verticillasters 2–13, cymes 1–4-flowered, 1 per node; proximal bracts oblanceolate, 2–10 × 0.5–1.5 mm; peduncles and pedicels spreading to ascending, retrorsely hairy with appressed, white, scalelike hairs, and sparsely glandular-pubescent. Flowers: calyx lobes ovate, 3.5–5.5 × 1.5–2 mm, scarious-margined, retrorsely hairy with appressed, white, scalelike hairs, and sparsely glandular-pubescent; corolla purple to violet, lined internally abaxially with reddish purple nectar guides, tubular-funnelform, 14–18 mm, sparsely white-lanate internally abaxially, tube 5–6 mm, throat gradually inflated, 4.5–6 mm diam., 2-ridged abaxially; stamens included, filaments glabrous or sparsely puberulent and sometimes with appressed, white, scalelike hairs proximally, pollen-sacs opposite, navicular, 0.9–1.4 mm, sutures papillate; staminode 7–9 mm, reaching orifice or exserted, flattened distally, 0.3–0.4 mm diam., tip straight to recurved, distal 4–5 mm pilose, hairs yellow, to 0.7 mm; style 9–11 mm. Capsules 5–7 × 4–5 mm.


Phenology: Flowering May–Jul(–Aug).
Habitat: Sandy or gravelly soils, pine-juniper woodlands, pine forests.
Elevation: 1200–2300 m.

Distribution

Calif., Mexico (Baja California)

Discussion

In California, Penstemon californicus is known from the San Jacinto Mountains in Orange and Riverside counties; it also occurs in the Sierra San Pedro Mártir in Baja California. The chromosome number of 2n = 16 reported by N. H. Holmgren (1993) is likely an error based on a count for Scrophularia californica reported in P. H. Raven et al. (1965) and later misattributed to P. californicus by D. V. Clark (1971) as n = 48 (the count listed in Raven et al.), and then corrected to 2n = 16.

Selected References

None.

Lower Taxa

None.

"/4+timescorollathroat" is not declared as a valid unit of measurement for this property.

... more about "Penstemon californicus"
mucronate +
Craig C. Freeman +
(Munz & I. M. Johnston) D. D. Keck +
alternate +, opposite +, whorled +, helical +  and subopposite +
tapered +
Penstemon linarioides var. californicus +
oblanceolate +
not +  and leathery +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
scarious-margined +
3.5mm;5.5mm +
glandular-pubescent +  and hairy +
short-tubular +  and short-campanulate +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
California beardtongue +
purple +  and violet +
white-lanate +
tubular-funnelform +
ampliate +  and ventricose-ampliate +
1.4 cm14 mm <br />0.014 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
1-4-flowered +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
scale-like +  and not needlelike +
Calif. +  and Mexico (Baja California) +
drupe-like +
1200–2300 m. +
curved +  and straight +
puberulent +  and glabrous +
1 +  and 4 +
Sandy or gravelly soils, pine-juniper woodlands, pine forests. +
yellow +  and white +
scale-like +
0 cm0 mm <br />0 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
membranous +
terminal +  and axillary +
persistent +  and deciduous +
cauline +, proximal +  and basal +
not +  and leathery +
orbiculate;linear;orbiculate;linear;truncate;oblanceolate;obovate;oblong;elliptic;round;lanceolate;ovate +
erose +  and entire +
glabrate +, hairy +  and glandular-pubescent +
toothed +  and entire +
herbaceous +  and scarious +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
spreading +  and ascending +
glandular-pubescent +  and hairy +
spreading;ascending +
glandular-pubescent +  and hairy +
5 +  and 4 +
Flowering May–Jul(–Aug). +
navicular-sigmoid +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Bull. Torrey Bot. Club +
minute +
dark-brown +  and black +
angled +  and rounded +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.21 cm2.1 mm <br />0.0021 m <br />) +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (?) +  and 0.9 cm9 mm <br />0.009 m <br /> (?) +
subalternate +, alternate +, whorled +  and opposite +
5 cm50 mm <br />0.05 m <br /> (8 cm80 mm <br />0.08 m <br />) +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
ascending;erect +
8 cm80 mm <br />0.08 m <br /> (30 cm300 mm <br />0.3 m <br />) +
0.9 cm9 mm <br />0.009 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
papillate +
denticulate +
Penstemon californicus +
Penstemon sect. Caespitosi +
species +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
2-ridged +  and inflated +
not constricted +
secund +, continuous +  and interrupted +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
cylindric +
2 cm20 mm <br />0.02 m <br /> (9 cm90 mm <br />0.09 m <br />) +
straight +
gibbous +  and not spurred +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
subshrub +  and cespitose +