Penstemon grinnellii

Eastwood

Bull. Torrey Bot. Club 32: 207. 1905.

Common names: Grinnell’s beardtongue
Endemic
Synonyms: Penstemon grinnellii subsp. scrophularioides (M. E. Jones) Munz P. grinnellii var. scrophularioides (M. E. Jones) N. H. Holmgren P. scrophularioides M. E. Jones
Treatment appears in FNA Volume 17. Treatment on page 251. Mentioned on page 152, 247.

Herbs. Stems ascending to erect, 10–90 cm, glaucous or not. Leaves: basal and proximal cauline 31–90 × 11–20 mm, blade oblanceolate to elliptic, base tapered, margins finely to coarsely dentate, apex rounded to obtuse; cauline 3–6 pairs, short-petiolate or sessile, (15–) 30–90 × 4–30 mm, blade lanceolate to ovate-oblong, base tapered, margins coarsely dentate, distals sometimes entire, apex obtuse or acute. Thyrses interrupted, cylindric, (5–) 7–34 cm, axis sparsely to densely glandular-pubescent, verticillasters 4–10, cymes 2–4-flowered; proximal bracts ovate to lanceolate, 5–50 × 2–18 mm; peduncles and pedicels spreading to ascending, glandular-pubescent. Flowers: calyx lobes ovate, 4.5–8.5 × (2–) 2.9–3.2 mm, glandular-pubescent; corolla white to pink or light lavender to light blue or light violet, with reddish purple nectar guides, strongly bilabiate, ventricose-ampliate, 22–35 mm, glandular-pubescent externally, glandular-pubescent internally, also white-villous abaxially, tube 5–8 mm, length 0.9–1.2 times calyx lobes, throat abruptly inflated, constricted at orifice, 10–18 mm diam., rounded abaxially; stamens included, filaments of shorter pair glandular-puberulent proximally, pollen-sacs navicular to subexplanate, 1.6–2.2 mm, sutures smooth; staminode 20–22 mm, exserted, 0.4–0.8 mm diam., tip recurved, distal 8–10 mm densely lanate, hairs yellowish or whitish, to 4 mm; proximal 6–8 mm densely glandular-pubescent; style 15–18 mm, glabrous. Capsules 10–14 × 5–8 mm, glabrous. 2n = 16.


Phenology: Flowering Apr–Jul.
Habitat: Chaparral, foothills, pine-juniper woodlands, pine forests.
Elevation: 500–2700 m.

Discussion

Two varieties of Penstemon grinnellii have been recognized. Variety grinnellii, with non-glaucous stems 10–60 cm and corollas white to pink and 22–30 mm, has been reported from the Western Transverse and Peninsular ranges, and in the San Bernardino, San Gabriel, and San Jacinto mountains. Variety scrophularioides, with glaucous stems 45–90 cm and corollas light lavender, light blue, or light violet and 26–35 mm, has been reported in the San Francisco Bay Area, southern Sierra Nevada and South Coast and Western Transverse ranges. Characteristics that distinguish the varieties overlap in some populations.

Penstemon ×peirsonii Munz & I. M. Johnston, a putative hybrid between P. grinnellii and P. speciosus, and P. ×dubius Davidson, a putative hybrid between P. centranthifolius and P. grinnellii, both have been reported from California (D. D. Keck 1937b).

Selected References

None.

Lower Taxa

None.

"/4+timescorollathroat" is not declared as a valid unit of measurement for this property."-1.2timescalyxlobes" is not declared as a valid unit of measurement for this property.

... more about "Penstemon grinnellii"
acute +, obtuse +  and rounded +
Craig C. Freeman +
Eastwood +
glandular-pubescent +
alternate +, opposite +, whorled +, helical +  and subopposite +
tapered;tapered +
lanceolate;ovate-oblong +
not +  and leathery +
0.2 cm2 mm <br />0.002 m <br /> (0.29 cm2.9 mm <br />0.0029 m <br />) +
4.5mm;8.5mm +
glandular-pubescent +
2.9mm;3.2mm +
short-tubular +  and short-campanulate +
1 cm10 mm <br />0.01 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
sessile +  and short-petiolate +
1.5 cm15 mm <br />0.015 m <br /> (3 cm30 mm <br />0.03 m <br />) +
3 cm30 mm <br />0.03 m <br /> (9 cm90 mm <br />0.09 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (3 cm30 mm <br />0.03 m <br />) +
Grinnell’s beardtongue +
white +  and pink or light lavender +
white-villous +  and glandular-pubescent +
cylindric-funnelform +, tubular-funnelform +  and ventricose +
ventricose-ampliate +
2.2 cm22 mm <br />0.022 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
2-4-flowered +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
scale-like +  and not needlelike +
drupe-like +
500–2700 m. +
curved +  and straight +
glandular-puberulent +
1 +  and 4 +
Chaparral, foothills, pine-juniper woodlands, pine forests. +
whitish +  and yellowish +
0 cm0 mm <br />0 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
membranous +
terminal +  and axillary +
persistent +  and deciduous +
3.1 cm31 mm <br />0.031 m <br /> (9 cm90 mm <br />0.09 m <br />) +
cauline +, proximal +  and basal +
glaucous +  and glabrous +
leathery +
1.1 cm11 mm <br />0.011 m <br /> (2 cm20 mm <br />0.02 m <br />) +
orbiculate;linear;orbiculate;linear;truncate;oblanceolate;obovate;oblong;elliptic;round;lanceolate;ovate +
erose +  and entire +
glandular-pubescent +  and glabrous +
denticulate +, toothed +  and entire +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
spreading +  and ascending +
glandular-pubescent +
spreading;ascending +
glandular-pubescent +
5 +  and 4 +
Flowering Apr–Jul. +
divergent +  and opposite +
navicular-sigmoid +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (5 cm50 mm <br />0.05 m <br />) +
ovate +  and lanceolate +
0.2 cm2 mm <br />0.002 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
glandular-pubescent +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Bull. Torrey Bot. Club +
minute +
brown +  and dark-brown or black +
rounded +  and angled +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
2 cm20 mm <br />0.02 m <br /> (?) +  and 2.2 cm22 mm <br />0.022 m <br /> (?) +
subalternate +, alternate +, whorled +  and opposite +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
ascending;erect +
not +  and glaucous +
10 cm100 mm <br />0.1 m <br /> (90 cm900 mm <br />0.9 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
papillate +
denticulate +
Penstemon grinnellii subsp. scrophularioides +, P. grinnellii var. scrophularioides +  and P. scrophularioides +
Penstemon grinnellii +
Penstemon sect. Spectabiles +
species +
1 cm10 mm <br />0.01 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
rounded +  and inflated +
constricted +
5 cm50 mm <br />0.05 m <br /> (7 cm70 mm <br />0.07 m <br />) +
cylindric +
7 cm70 mm <br />0.07 m <br /> (34 cm340 mm <br />0.34 m <br />) +
straight +
0 cm0 mm <br />0 m <br /> (0.01 cm0.1 mm <br />1.0e-4 m <br />) +
gibbous +  and not spurred +
0.5 cm5 mm <br />0.005 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +