Penstemon harbourii

A. Gray

Proc. Amer. Acad. Arts 6: 71. 1862.

Common names: Harbour’s beardtongue
Endemic
Treatment appears in FNA Volume 17. Treatment on page 206. Mentioned on page 188, 207.

Herbs. Caudex rhizomelike. Stems decumbent, ascending, or weakly erect, 4–18 cm, retrorsely hairy, not glaucous. Leaves cauline, not leathery, glabrate, retrorsely hairy, or puberulent; cauline 2–4 pairs, petiolate or sessile, 7–28 × 3–12 mm, blade spatulate to oblanceolate, base tapered to clasping, margins entire, apex rounded or obtuse to acute. Thyrses interrupted or continuous, ± secund, 1–3 cm, axis ± glandular-pubescent, verticillasters 1 or 2 (or 3), cymes 1-flowered or 2-flowered, 1 or 2 per node; proximal bracts lanceolate, 17–23 × 4–10 mm, margins entire; peduncles and pedicels ascending to erect, retrorsely hairy and glandular-pubescent. Flowers: calyx lobes lanceolate, 6–10 × 1.3–2 mm, glandular-pubescent; corolla lavender or bluish lavender to lavender-purple, without nectar guides, bilabiate, not personate, tubular-funnelform, 15–20 mm, glandular-pubescent externally, densely white-lanate internally abaxially, tube 4–6 mm, throat gradually inflated, 5–6 mm diam., prominently 2-ridged abaxially; stamens included, pollen-sacs opposite, subexplanate, 0.7–0.9 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 8–9 mm, included, 0.3–0.4 mm diam., tip recurved, distal 4–5 mm moderately to densely lanate, hairs yellow, to 1 mm; style 11–12 mm. Capsules 6–8 × 4–5 mm, glabrous. 2n = 16.


Phenology: Flowering Jun–Aug.
Habitat: Alpine talus, gravel slopes, boulder fields.
Elevation: 3200–4200 m.

Discussion

Penstemon harbourii is found on the high peaks of the Central Rocky Mountains. Populations have been documented in at least 17 counties. Habit, pubescence, and root characters of P. harbourii are somewhat anomalous in sect. Penstemon, prompting D. D. Keck (1945) to suggest possible ties with sect. Caespitosi or sect. Cristati. Molecular data appear to provide support for ties with sect. Caespitosi (A. D. Wolfe et al. 2006; C. A. Wessinger et al. 2016).

Selected References

None.

Lower Taxa

None.

"/4+timescorollathroat" is not declared as a valid unit of measurement for this property.

... more about "Penstemon harbourii"
obtuse +  and acute +
Craig C. Freeman +
A. Gray +
glandular-pubescent +
alternate +, subopposite +, whorled +  and opposite +
glaucescent +
alternate +, opposite +, whorled +, helical +  and subopposite +
cauline +, proximal +  and basal +
rarely +, not glaucous +, glabrate +, pubescent +, hairy +, glandular-lanate +, glandular-pubescent +, puberulent +, glandular +  and glabrous +
not +  and leathery +
tapered +
spatulate;oblanceolate +
not +  and leathery +
glandular-pubescent +
short-tubular +  and short-campanulate +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
sessile +  and petiolate +
0.7 cm7 mm <br />0.007 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
Harbour’s beardtongue +
not personate +  and bilabiate +
yellow +, purple +, violet +, blue +, lavender +, pink +, ochroleucous +  and white +
bluish lavender to lavender-purple +  and lavender +
white-lanate +  and glandular-pubescent +
tubular-funnelform +
ampliate +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
2-flowered +  and 1-flowered +
2 +  and 1 +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
scale-like +  and not needlelike +
drupe-like +
3200–4200 m. +
curved +  and straight +
1 +  and 4 +
Alpine talus, gravel slopes, boulder fields. +
0 cm0 mm <br />0 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
membranous +
terminal +  and axillary +
persistent +  and deciduous +
puberulent +, hairy +  and glabrate +
not leathery +
orbiculate;linear;orbiculate;linear;truncate;oblanceolate;obovate;oblong;elliptic;round;lanceolate;ovate +
glutinous +
glandular-ciliolate +, hairy +, puberulent +, glabrescent +, glabrous +  and glandular-pubescent +
lacerate +, toothed +  and entire +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
ascending +  and erect +
glandular-pubescent +  and hairy +
ascending;erect +
glandular-pubescent +  and hairy +
5 +  and 4 +
Flowering Jun–Aug. +
navicular-sigmoid +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
1/5 +  and 2/3 +
1.7 cm17 mm <br />0.017 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Proc. Amer. Acad. Arts +
minute +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
black +, gray +, brown +  and tan +
angled +  and slightly rounded +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (?) +  and 0.9 cm9 mm <br />0.009 m <br /> (?) +
subalternate +, alternate +, whorled +  and opposite +
glutinous +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
erect;ascending;erect;ascending;decumbent +
not glaucous +  and hairy +
4 cm40 mm <br />0.04 m <br /> (18 cm180 mm <br />0.18 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
papillate +
denticulate +
Penstemon harbourii +
Penstemon sect. Penstemon +
species +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
2-ridged +  and inflated +
not constricted +
secund +, continuous +  and interrupted +
cylindric +  and conic +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
straight +
gibbous +  and not spurred +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +