Penstemon heterophyllus

Lindley

Edwards’s Bot. Reg. 22: plate 1899. 1836.

Common names: Foothill beardtongue
Endemic
Treatment appears in FNA Volume 17. Treatment on page 235. Mentioned on page 230, 236.

Subshrubs. Stems ascending to erect, 30–150 cm, glabrous or retrorsely hairy, hairs pointed, rarely glaucous. Leaves essentially cauline, basal absent or essentially so, opposite, glabrous or retrorsely hairy to puberulent, hairs pointed, rarely glaucous; cauline 5–11 pairs, petiolate or sessile, 10–100 × 0.5–12 mm, blade oblanceolate to lanceolate or linear, base tapered to clasping, margins entire, apex acute. Thyrses interrupted or continuous, cylindric, (6–) 10–32 cm, axis glabrous or retrorsely hairy, verticillasters (3–) 5–14, cymes 1 (–3) -flowered, (1 or) 2 per node; proximal bracts lanceolate to linear, 5–65 × 0.5–5 mm; peduncles and pedicels ascending to erect, glabrous or retrorsely hairy. Flowers: calyx lobes obovate to oblanceolate or ovate to lanceolate, 2.6–6 (–7) × 1.3–3.3 mm, glabrous or puberulent; corolla blue to violet or purple, without nectar guides, ventricose to ventricose-ampliate, 24–38 mm, glabrous externally, glabrous internally, tube 7–10 mm, throat gradually inflated, 8–11 mm diam., 2-ridged abaxially; stamens included or longer pair reaching orifice or slightly exserted, filaments glabrous, pollen-sacs parallel, 2.2–3 mm, distal 1/3–1/2 indehiscent, sides pubescent to lanate, hairs white, to 0.8 mm near filament attachment, sutures denticulate, teeth to 0.2 mm; staminode 16–19 mm, included, 0.4–0.6 mm diam., glabrous; style 18–27 mm. Capsules 6–9 × 4–5 mm. 2n = 16, 32.

Discussion

Varieties 3 (3 in the flora).

Penstemon heterophyllus apparently occurs only in California (see discussion under 209c. var. purdyi). The species includes three varieties concentrated in the Coast Ranges from Humboldt County south to San Bernardino and San Diego counties.

Selected References

None.

Key

1 Thyrse axes glabrous; stems glabrous, rarely retrorsely hairy; calyx lobes obovate to oblanceolate, rarely lanceolate. Penstemon heterophyllus var. heterophyllus
1 Thyrse axes retrorsely hairy; stems retrorsely hairy; calyx lobes ovate to lanceolate. > 2
2 Cauline leaves 15–45 × 0.5–4 mm, axillary fascicles present, sometimes absent. Penstemon heterophyllus var. australis
2 Cauline leaves (25–)30–100 × 2–12 mm, axillary fascicles absent, rarely present. Penstemon heterophyllus var. purdyi

"/4+timescorollathroat" is not declared as a valid unit of measurement for this property.

... more about "Penstemon heterophyllus"
Craig C. Freeman +
Lindley +
hairy +  and glabrous +
subalternate +, whorled +, alternate +  and opposite +
cauline +, proximal +  and basal +
not +, glaucous +, glandular-pubescent +, hirsute +, pilose +, canescent +, hairy +, puberulent +, glabrate +  and glabrous +
tapered +
oblanceolate;lanceolate or linear +
not +  and leathery +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
puberulent +  and glabrous +
obovate;oblanceolate or ovate +
1.3mm;3.3mm +
short-tubular +  and short-campanulate +
0.6 cm6 mm <br />0.006 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
sessile +  and petiolate +
1 cm10 mm <br />0.01 m <br /> (10 cm100 mm <br />0.1 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
Foothill beardtongue +
not personate +  and bilabiate +
blue +  and violet or purple +
ventricose +
ventricose-ampliate +
2.4 cm24 mm <br />0.024 m <br /> (3.8 cm38 mm <br />0.038 m <br />) +
1(-3)-flowered +
scale-like +  and not needlelike +
drupe-like +
curved +  and straight +
1 +  and 4 +
0 cm0 mm <br />0 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
pointed +
0 cm0 mm <br />0 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
membranous +
terminal +  and axillary +
persistent +  and deciduous +
orbiculate;linear;orbiculate;linear;truncate;oblanceolate;obovate;oblong;elliptic;round;lanceolate;ovate +
erose +  and entire +
ciliolate +, pubescent +, puberulent +, scabrous +, glandular-pubescent +  and glabrous +
lacerate +, laciniate-pinnatifid +, toothed +  and entire +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
ascending +  and erect +
hairy +  and glabrous +
ascending;erect +
hairy +  and glabrous +
5 +  and 4 +
navicular-sigmoid +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
1/3 +  and 1/2 +
0.5 cm5 mm <br />0.005 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
lanceolate +  and linear +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Edwards’s Bot. Reg. +
minute +
brown +, gray +  and tan +
patelliform +  and angled +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.34 cm3.4 mm <br />0.0034 m <br />) +
pubescent +  and lanate +
exserted +  and included +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
1.6 cm16 mm <br />0.016 m <br /> (?) +  and 1.9 cm19 mm <br />0.019 m <br /> (?) +
subalternate +, alternate +, whorled +  and opposite +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
ascending;erect +
hairy +  and glabrous +
30 cm300 mm <br />0.3 m <br /> (150 cm1,500 mm <br />1.5 m <br />) +
1.8 cm18 mm <br />0.018 m <br /> (2.7 cm27 mm <br />0.027 m <br />) +
papillate +
denticulate +
Penstemon subg. Saccanthera +
Penstemon heterophyllus +
Penstemon sect. Saccanthera +
species +
0.8 cm8 mm <br />0.008 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
2-ridged +  and inflated +
not constricted +
continuous +  and interrupted +
6 cm60 mm <br />0.06 m <br /> (10 cm100 mm <br />0.1 m <br />) +
cylindric +
10 cm100 mm <br />0.1 m <br /> (32 cm320 mm <br />0.32 m <br />) +
straight +
0 cm0 mm <br />0 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
gibbous +  and not spurred +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +