Penstemon humilis var. humilis

IllustratedEndemic
Synonyms: Penstemon cinereus Piper P. cinereus subsp. foliatus D. D. Keck P. decurvus Pennell ex Crosswhite P. humilis var. desereticus S. L. Welsh
Treatment appears in FNA Volume 17. Treatment on page 209. Mentioned on page 201.

Leaves: basal leaves puberulent or retrorsely hairy, (8–) 15–105 × 2–13 (–18) mm, cauline blade margins entire, rarely ± serrate distally. Flowers: corolla (7–) 12–19 mm. 2n = 16.


Phenology: Flowering May–Jul.
Habitat: Open, rocky slopes, hillsides, sagebrush shrublands, pine-juniper woodlands, coniferous forests, alpine meadows.
Elevation: 1000–3200 m.

Distribution

Calif., Colo., Idaho, Mont., Nev., Oreg., Utah, Wash., Wyo.

Discussion

Welsh described var. desereticus based on material with relatively larger corollas (15–19 mm) and smaller basal leaves (8–25 × 2–6 mm) from mountain ranges in the western Bonneville Basin of Utah; E. C. Neese and N. D. Atwood (2003) considered the variety to be limited to Utah. Variety desereticus appears to be confluent with a form of the species in the Calcareous Mountains of eastern Nevada, which N. H. Holmgren (1984) discussed and later recognized as Penstemon decurvus (Holmgren 2017). Penstemon decurvus, described from eastern Lincoln County, Nevada, also is referable here. Plants from northeastern California, extreme southwestern Idaho, extreme northwestern Nevada, and eastern and central Oregon often have cinereous leaves and relatively longer stems, shorter calyx lobes, and, sometimes, more open inflorescences; these plants have been called P. cinereus. This element grades into other phases to the east and appears to be another form of highly variable var. humilis. Some specimens from eastern Idaho (Clark and Fremont counties) have glabrous leaves, possibly from genetic exchange with P. aridus.

Selected References

None.

Lower Taxa

None.

"/4+timescorollathroat" is not declared as a valid unit of measurement for this property.

glandular-pubescent +
acute +  and obtuse +
Craig C. Freeman +
Nuttall ex A. Gray +
glandular-pubescent +
alternate +, subopposite +, whorled +  and opposite +
glaucescent +
alternate +, opposite +, whorled +, helical +  and subopposite +
0.8 cm8 mm <br />0.008 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
1.3 cm13 mm <br />0.013 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (10.5 cm105 mm <br />0.105 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
cauline +, proximal +  and basal +
rarely +, not glaucous +, glabrate +, pubescent +, hairy +, glandular-lanate +, glandular-pubescent +, puberulent +, glandular +  and glabrous +
not +  and leathery +
tapered;tapered +
obovate;oblanceolate or lanceolate +
not +  and leathery +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
ovate;lanceolate +
short-tubular +  and short-campanulate +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
petiolate +  and sessile +
not personate +  and bilabiate +
0.7 cm7 mm <br />0.007 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
white +, blue +  and purple or violet +
yellow-lanate +  and glandular-pubescent +
tubular-funnelform +
ampliate +
1.2 cm12 mm <br />0.012 m <br /> (1.9 cm19 mm <br />0.019 m <br />) +
2-6(-11)-flowered +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
scale-like +  and not needlelike +
Calif. +, Colo. +, Idaho +, Mont. +, Nev. +, Oreg. +, Utah +, Wash. +  and Wyo. +
drupe-like +
1000–3200 m. +
curved +  and straight +
1 +  and 4 +
Open, rocky slopes, hillsides, sagebrush shrublands, pine-juniper woodlands, coniferous forests, alpine meadows. +
golden yellow +  and yellow +
0 cm0 mm <br />0 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
membranous +
terminal +  and axillary +
persistent +  and deciduous +
basal +  and cauline +
hairy +  and glabrous +
not leathery +
orbiculate;linear;orbiculate;linear;truncate;oblanceolate;obovate;oblong;elliptic;round;lanceolate;ovate +
erose +  and entire +
entire +, serrate +  and serrulate +
glutinous +
glandular-ciliolate +, hairy +, puberulent +, glabrescent +, glabrous +  and glandular-pubescent +
lacerate +, toothed +  and entire +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
ascending +  and erect +
hairy +  and glandular-pubescent +
ascending;erect +
hairy +  and glandular-pubescent +
5 +  and 4 +
Flowering May–Jul. +
navicular-sigmoid +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
1/5 +  and 2/3 +
short-petiolate +
3.5 cm35 mm <br />0.035 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
3.4 cm34 mm <br />0.034 m <br /> (4.8 cm48 mm <br />0.048 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (3.4 cm34 mm <br />0.034 m <br />) +
ovate +  and lanceolate +
0.1 cm1 mm <br />0.001 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
1 cm10 mm <br />0.01 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
Proc. Amer. Acad. Arts +
minute +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
black +, gray +, brown +  and tan +
angled +  and slightly rounded +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Illustrated +  and Endemic +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (?) +  and 0.9 cm9 mm <br />0.009 m <br /> (?) +
subalternate +, alternate +, whorled +  and opposite +
30 cm300 mm <br />0.3 m <br /> (65 cm650 mm <br />0.65 m <br />) +
glutinous +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
decumbent;erect +
not glaucous +  and hairy +
5 cm50 mm <br />0.05 m <br /> (30 cm300 mm <br />0.3 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
papillate +  and smooth +
denticulate +
Penstemon cinereus +, P. cinereus subsp. foliatus +, P. decurvus +  and P. humilis var. desereticus +
Penstemon humilis var. humilis +
Penstemon humilis +
variety +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
2-ridged +  and inflated +
not constricted +
secund +  and interrupted +
20 cm200 mm <br />0.2 m <br /> (39 cm390 mm <br />0.39 m <br />) +
cylindric +
3 cm30 mm <br />0.03 m <br /> (20 cm200 mm <br />0.2 m <br />) +
straight +
gibbous +  and not spurred +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
cauline +  and proximal +