Penstemon leiophyllus

Pennell

Contr. U.S. Natl. Herb. 20: 346. 1920.

Common names: Charleston beardtongue
Endemic
Treatment appears in FNA Volume 17. Treatment on page 170. Mentioned on page 156, 171.

Stems ascending to erect, 4–55 (–80) cm, glabrous, not glaucous. Leaves basal and cauline, not leathery, glabrous, not glaucous; basal and proximal cauline (20–) 30–100 × 4–15 (–20) mm, blade oblanceolate, base tapered, margins entire, apex obtuse to acute; cauline 2–7 pairs, sessile or proximals sometimes short-petiolate, 16–80 × 2–20 mm, blade lanceolate to linear, base tapered, apex obtuse to acute. Thyrses interrupted or continuous, secund, (1–) 3–30 (–40) cm, axis glandular-pubescent, verticillasters 2–8 (–12), cymes 1–3 (or 4) -flowered, (1 or) 2 per node; proximal bracts lanceolate, (6–) 10–60 × (1–) 3–24 mm; peduncles and pedicels glandular-pubescent. Flowers: calyx lobes lanceolate, (3.8–) 4.5–8.5 × 1.6–3.8 mm, glandular-pubescent; corolla violet to blue or purple, with violet or reddish purple nectar guides, ventricose, 18–35 mm, glandular-pubescent externally, glabrous or white-lanate internally abaxially, glandular-pubescent adaxially, tube 6–10 mm, throat gradually inflated, not constricted at orifice, 5–10 mm diam., 2-ridged abaxially; stamens: longer pair reaching orifice, pollen-sacs divergent to nearly opposite, navicular, 1.2–2 mm, dehiscing incompletely, proximal 1/5 indehiscent, connective not splitting, sides glabrous, sutures denticulate, teeth to 0.1 mm; staminode 10–14 mm, included, 0.4–0.5 mm diam., tip straight, distal 1–5 mm sparsely to densely pilose, hairs yellow or orangish yellow, to 1 mm; style 12–19 mm. Capsules 7–11 × 5–6 mm.

Discussion

Varieties 3 (3 in the flora).

F. S. Crosswhite (1967) recognized three species in the Penstemon leiophyllus complex: P. francisci-pennellii, P. keckii, and P. leiophyllus. N. H. Holmgren (1984) expanded P. leiophyllus to include the other species as varieties. The complex is unique in the section in having patches of glandular hairs on the internal, adaxial surfaces of the corolla throats. The varieties are essentially allopatric; characters distinguishing them are minor.

Selected References

None.

Key

1 Stems 15–55(–80) cm; cauline leaves 4–20 mm wide, blades lanceolate, usually flat and straight; corollas glabrous, rarely white-lanate, internally abaxially; Utah. Penstemon leiophyllus var. leiophyllus
1 Stems 4–25(–43) cm; cauline leaves 2–5(–10) mm wide, blades lanceolate to linear, usually folded lengthwise and curved; corollas white-lanate, rarely glabrous, internally abaxially; Nevada. > 2
2 Corollas (22–)25–35 mm; Lincoln, Nye, and White Pine counties, Nevada. Penstemon leiophyllus var. francisci-pennellii
2 Corollas 18–22 mm; Clark County, Nevada. Penstemon leiophyllus var. keckii

"/4+timescorollathroat" is not declared as a valid unit of measurement for this property.

... more about "Penstemon leiophyllus"
obtuse +  and acute +
Craig C. Freeman +
Pennell +
glandular-pubescent +
alternate +, opposite +, whorled +, helical +  and subopposite +
tapered;tapered +
Penstemon +
lanceolate;linear +
not +  and leathery +
0.38 cm3.8 mm <br />0.0038 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
4.5mm;8.5mm +
glandular-pubescent +
1.6mm;3.8mm +
short-tubular +  and short-campanulate +
0.7 cm7 mm <br />0.007 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Charleston beardtongue +
not personate +, weakly bilabiate +  and strongly bilabiate +
violet +  and blue or purple +
glandular-pubescent +, white-lanate +  and glabrous +
ventricose +
ventricose-ampliate +
1.8 cm18 mm <br />0.018 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
1-3-flowered +
scale-like +  and not needlelike +
Nev. +  and Utah +
drupe-like +
curved +  and straight +
1 +  and 4 +
orangish yellow +  and yellow +
0 cm0 mm <br />0 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
membranous +
terminal +  and axillary +
persistent +  and deciduous +
basal +  and cauline +
not glaucous +  and glabrous +
not leathery +
orbiculate;linear;orbiculate;linear;truncate;oblanceolate;obovate;oblong;elliptic;round;lanceolate;ovate +
glutinous +
puberulent +, glandular-pubescent +  and glabrous +
lacerate +
reddish purple +  and violet +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
0 cm0 mm <br />0 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
spreading +, erect +  and ascending +
glandular-pubescent +
spreading;erect;ascending +
glandular-pubescent +
5 +  and 4 +
divergent +  and nearly opposite +
sigmoid +  and navicular-sigmoid +
indehiscent +  and dehiscing +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
short-petiolate +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
1 cm10 mm <br />0.01 m <br /> (6 cm60 mm <br />0.06 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (2.4 cm24 mm <br />0.024 m <br />) +
1.6 cm16 mm <br />0.016 m <br /> (8 cm80 mm <br />0.08 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Contr. U.S. Natl. Herb. +
minute +
tan +  and brown or dark-brown +
angled +  and angled-elongate +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
papillate +
exserted +  and included +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
1 cm10 mm <br />0.01 m <br /> (?) +  and 1.4 cm14 mm <br />0.014 m <br /> (?) +
short-petiolate +  and petiolate +
55 cm550 mm <br />0.55 m <br /> (80 cm800 mm <br />0.8 m <br />) +
glutinous +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
ascending;erect +
cauline +, proximal +  and basal +
not glaucous +  and glabrous +
reduced +
4 cm40 mm <br />0.04 m <br /> (55 cm550 mm <br />0.55 m <br />) +
not +  and leathery +
1.2 cm12 mm <br />0.012 m <br /> (1.9 cm19 mm <br />0.019 m <br />) +
papillate +
denticulate +
Penstemon subg. Habroanthus +
Penstemon leiophyllus +
Penstemon sect. Glabri +
species +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
2-ridged +  and inflated +
not constricted +
secund +, continuous +  and interrupted +
30 cm300 mm <br />0.3 m <br /> (40 cm400 mm <br />0.4 m <br />) +
cylindric +
3 cm30 mm <br />0.03 m <br /> (30 cm300 mm <br />0.3 m <br />) +
straight +
0.1 cm1 mm <br />0.001 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
smooth +  and papillate +
0 cm0 mm <br />0 m <br /> (0.01 cm0.1 mm <br />1.0e-4 m <br />) +
gibbous +  and not spurred +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
petiolate +  and sessile +
cauline +  and proximal +