Penstemon neotericus

D. D. Keck

Univ. Calif. Publ. Bot. 16: 398, fig. 13. 1932.

Common names: Plumas County beardtongue
Endemic
Treatment appears in FNA Volume 17. Treatment on page 238. Mentioned on page 231, 239.

Subshrubs. Stems ascending to erect, 20–80 cm, glabrous, sometimes ± puberulent, hairs pointed, glaucous. Leaves basal and cauline, basal sometimes few, opposite, glabrous, sometimes proximals puberulent, hairs pointed, glaucous; basal and proximal cauline 15–80 × 2–10 mm, blade spatulate to oblanceolate, base tapered, margins entire, apex rounded to obtuse or acute; cauline 4–6 pairs, sessile, 20–80 × 3–9 mm, blade ovate to lanceolate, base tapered, margins entire, apex acute. Thyrses interrupted, cylindric, 5–25 cm, axis glandular-pubescent, verticillasters 4–6, cymes 1–3-flowered, 2 per node; proximal bracts ovate to lanceolate, 4–50 × 1–8 mm; peduncles and pedicels ascending, glandular-pubescent. Flowers: calyx lobes ovate to lanceolate, 3.8–6 × 1.5–2.2 mm, glandular-pubescent; corolla violet to blue or purple, without nectar guides, ventricose to ventricose-ampliate, 25–38 mm, glandular-pubescent externally, glabrous internally, tube 7–9 mm, throat gradually inflated, 8–10 mm diam., 2-ridged abaxially; stamens included, filaments glabrous, pollen-sacs parallel, 2.4–3.2 mm, distal 1/5–1/3 indehiscent, sides pilose to lanate, hairs white, to 0.6 mm near filament attachment, sutures denticulate, teeth to 0.2 mm; staminode 11–15 mm, included, 0.4–0.6 mm diam., glabrous; style 15–20 mm. Capsules 7–9 × 4–5 mm. 2n = 64.


Phenology: Flowering May–Aug(–Oct).
Habitat: Volcanic soils, scrub, open pine and fir forests.
Elevation: 1000–2200 m.

Discussion

Penstemon neotericus is known from the Cascade Range and northern Sierra Nevada of northern California (Butte, Lassen, Placer, Plumas, Shasta, Sierra, Siskiyou, Tehama, and Yuba counties). D. D. Keck (1932) used morphologic and geographic data to hypothesize that P. neotericus, an octoploid, arose through hybridization between P. azureus and P. laetus. J. Clausen (1933) provided cytological evidence supporting that hypothesis. A diploid chromosome number of 2n = 32 listed by N. H. Holmgren (1993) is probably a transcription error.

Lower Taxa

None.

"/4+timescorollathroat" is not declared as a valid unit of measurement for this property.

... more about "Penstemon neotericus"
acute +, rounded +  and obtuse or acute +
Craig C. Freeman +
D. D. Keck +
glandular-pubescent +
subalternate +, whorled +, alternate +  and opposite +
alternate +, opposite +, whorled +, helical +  and subopposite +
cauline +, proximal +  and basal +
not +, glaucous +, glandular-pubescent +, hirsute +, pilose +, canescent +, hairy +, puberulent +, glabrate +  and glabrous +
tapered;tapered +
ovate;lanceolate +
not +  and leathery +
glandular-pubescent +
ovate;lanceolate +
1.5mm;2.2mm +
short-tubular +  and short-campanulate +
0.7 cm7 mm <br />0.007 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
2 cm20 mm <br />0.02 m <br /> (8 cm80 mm <br />0.08 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
short-petiolate +, petiolate +  and sessile +
Plumas County beardtongue +
not personate +  and bilabiate +
violet +  and blue or purple +
glabrous +  and glandular-pubescent +
ventricose +
ventricose-ampliate +
2.5 cm25 mm <br />0.025 m <br /> (3.8 cm38 mm <br />0.038 m <br />) +
1-3-flowered +
scale-like +  and not needlelike +
drupe-like +
1000–2200 m. +
curved +  and straight +
1 +  and 4 +
Volcanic soils, scrub, open pine and fir forests. +
0 cm0 mm <br />0 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
pointed +
0 cm0 mm <br />0 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
membranous +
terminal +  and axillary +
persistent +  and deciduous +
cauline +  and basal +
orbiculate;linear;orbiculate;linear;truncate;oblanceolate;obovate;oblong;elliptic;round;lanceolate;ovate +
erose +  and entire +
ciliolate +, pubescent +, puberulent +, scabrous +, glandular-pubescent +  and glabrous +
lacerate +, laciniate-pinnatifid +, toothed +  and entire +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
glandular-pubescent +
glandular-pubescent +
5 +  and 4 +
Flowering May–Aug(–Oct). +
navicular-sigmoid +
0.24 cm2.4 mm <br />0.0024 m <br /> (0.32 cm3.2 mm <br />0.0032 m <br />) +
1/5 +  and 1/3 +
0.4 cm4 mm <br />0.004 m <br /> (5 cm50 mm <br />0.05 m <br />) +
ovate +  and lanceolate +
0.1 cm1 mm <br />0.001 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Univ. Calif. Publ. Bot. +
clausen1933a +
minute +
brown +, gray +  and tan +
patelliform +  and angled +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.34 cm3.4 mm <br />0.0034 m <br />) +
pilose +  and lanate +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (?) +  and 1.5 cm15 mm <br />0.015 m <br /> (?) +
subalternate +, alternate +, whorled +  and opposite +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
ascending;erect +
puberulent +  and glabrous +
20 cm200 mm <br />0.2 m <br /> (80 cm800 mm <br />0.8 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
papillate +
denticulate +
Penstemon subg. Saccanthera +
Penstemon neotericus +
Penstemon sect. Saccanthera +
species +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
2-ridged +  and inflated +
not constricted +
cylindric +
5 cm50 mm <br />0.05 m <br /> (25 cm250 mm <br />0.25 m <br />) +
straight +
0 cm0 mm <br />0 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
gibbous +  and not spurred +
0.7 cm7 mm <br />0.007 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
cauline +  and proximal +