Penstemon pennellianus

D. D. Keck

Amer. Midl. Naturalist 23: 614. 1940.

Common names: Pennell’s beardtongue
Endemic
Treatment appears in FNA Volume 17. Treatment on page 176. Mentioned on page 161.

Stems ascending to erect, 18–85 cm, glabrous, not glaucous. Leaves basal and cauline, ± leathery or not, glabrous, not glaucous; basal and proximal cauline 55–250 × 7–40 mm, blade lanceolate to elliptic, base tapered, margins entire, apex obtuse to acute; cauline 3–5 pairs, short-petiolate or sessile, 40–83 × 10–36 mm, blade ovate to lanceolate, base tapered to clasping, apex obtuse to acute. Thyrses continuous, cylindric, 4–35 cm, axis glabrous, verticillasters 4–11, cymes 1–6-flowered, 2 per node; proximal bracts ovate to lanceolate, 22–85 × 11–35 mm; peduncles and pedicels glabrous, peduncles to 20 mm, pedicels 1–6 mm. Flowers: calyx lobes ovate to lanceolate, (5.3–) 6.5–8.5 × 1.6–3 mm, apex acuminate to caudate, glabrous; corolla blue to violet or purple, with reddish purple nectar guides, ventricose, 29–35 mm, glabrous externally, glabrous internally, tube 8–11 mm, throat gradually inflated, not constricted at orifice, 9–10 mm diam., slightly 2-ridged abaxially; stamens: longer pair reaching orifice, pollen-sacs divergent to opposite, sigmoid, 2–2.7 mm, dehiscing incompletely, proximal 1/5–1/3 indehiscent, connective not splitting, sides sparsely hirsute, hairs white or tan, to 0.1 mm, especially near connective, sometimes merely scurfy near connective, sutures denticulate, teeth to 0.1 mm; staminode 16–20 mm, reaching orifice, 0.5–1 mm diam., tip straight, distal 4–6 mm ± uniformly sparsely pilose, hairs yellow, to 1 mm; style 20–24 mm. Capsules 10–13 × 8–10 mm.


Phenology: Flowering May–Jun.
Habitat: Rocky ridges, sandy to rocky slopes, coniferous forests.
Elevation: 1300–1800 m.

Discussion

Penstemon pennellianus occurs primarily in the Blue Mountains of northeastern Oregon (Grant, Umatilla, Union, and Wallowa counties) and southeastern Washington (Asotin, Columbia, and Garfield counties).

Selected References

None.

Lower Taxa

None.

"/4+timescorollathroat" is not declared as a valid unit of measurement for this property.

... more about "Penstemon pennellianus"
acuminate +  and caudate +
Craig C. Freeman +
D. D. Keck +
alternate +, opposite +, whorled +, helical +  and subopposite +
tapered;tapered +
Penstemon +
ovate;lanceolate +
not +  and leathery +
0.53 cm5.3 mm <br />0.0053 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
6.5mm;8.5mm +
ovate;lanceolate +
short-tubular +  and short-campanulate +
1 cm10 mm <br />0.01 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
sessile +  and short-petiolate +
4 cm40 mm <br />0.04 m <br /> (8.3 cm83 mm <br />0.083 m <br />) +
1 cm10 mm <br />0.01 m <br /> (3.6 cm36 mm <br />0.036 m <br />) +
Pennell’s beardtongue +
not personate +, weakly bilabiate +  and strongly bilabiate +
blue +  and violet +
ventricose +
ventricose-ampliate +
2.9 cm29 mm <br />0.029 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
1-6-flowered +
scale-like +  and not needlelike +
Oreg. +  and Wash. +
drupe-like +
1300–1800 m. +
curved +  and straight +
1 +  and 4 +
Rocky ridges, sandy to rocky slopes, coniferous forests. +
yellow +, tan +  and white +
0 cm0 mm <br />0 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
membranous +
terminal +  and axillary +
persistent +  and deciduous +
basal +  and cauline +
not glaucous +  and glabrous +
not +  and leathery +
orbiculate;linear;orbiculate;linear;truncate;oblanceolate;obovate;oblong;elliptic;round;lanceolate;ovate +
glutinous +
puberulent +, glandular-pubescent +  and glabrous +
lacerate +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
0 cm0 mm <br />0 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
spreading +, erect +  and ascending +
0.1 cm1 mm <br />0.001 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
spreading;erect;ascending +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
5 +  and 4 +
Flowering May–Jun. +
divergent +  and opposite +
indehiscent +  and dehiscing +
0.2 cm2 mm <br />0.002 m <br /> (0.27 cm2.7 mm <br />0.0027 m <br />) +
1/5 +  and 1/3 +
2.2 cm22 mm <br />0.022 m <br /> (8.5 cm85 mm <br />0.085 m <br />) +
ovate +  and lanceolate +
1.1 cm11 mm <br />0.011 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
Amer. Midl. Naturalist +
minute +
tan +  and brown or dark-brown +
angled +  and angled-elongate +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
papillate +
exserted +  and included +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
included +  and exserted +
1.6 cm16 mm <br />0.016 m <br /> (?) +  and 2 cm20 mm <br />0.02 m <br /> (?) +
short-petiolate +  and petiolate +
glutinous +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
ascending;erect +
cauline +, proximal +  and basal +
not glaucous +  and glabrous +
reduced +
18 cm180 mm <br />0.18 m <br /> (85 cm850 mm <br />0.85 m <br />) +
not +  and leathery +
2 cm20 mm <br />0.02 m <br /> (2.4 cm24 mm <br />0.024 m <br />) +
papillate +
denticulate +
Penstemon subg. Habroanthus +
Penstemon pennellianus +
Penstemon sect. Glabri +
species +
0.9 cm9 mm <br />0.009 m <br /> (1 cm10 mm <br />0.01 m <br />) +
2-ridged +  and inflated +
not constricted +
cylindric +
4 cm40 mm <br />0.04 m <br /> (35 cm350 mm <br />0.35 m <br />) +
straight +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
smooth +  and papillate +
0 cm0 mm <br />0 m <br /> (0.01 cm0.1 mm <br />1.0e-4 m <br />) +
gibbous +  and not spurred +
0.8 cm8 mm <br />0.008 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
petiolate +  and sessile +
cauline +  and proximal +