Penstemon sudans

M. E. Jones

Contr. W. Bot. 8: 37. 1898.

Common names: Susanville beardtongue
Endemic
Synonyms: Penstemon deustus subsp. sudans (M. E. Jones) Pennell & D. D. Keck
Treatment appears in FNA Volume 17. Treatment on page 223. Mentioned on page 183, 186.

Subshrubs. Stems ascending to erect, 18–70 cm, glandular-pubescent, sometimes densely so distally, not glaucous. Leaves cauline, opposite or subopposite, not leathery, glandular-pubescent; cauline (4–) 7–10 pairs, sessile or proximals short-petiolate, 14–60 × 8–20 (–30) mm, blade obovate to ovate or lanceolate, base tapered to clasping, margins dentate, dentate-serrate, or serrate, apex acute to acuminate. Thyrses interrupted or continuous, cylindric, 13–35 cm, axis densely glandular-pubescent, verticillasters 9–18, cymes 1–5-flowered, 2 per node; proximal bracts ovate to lanceolate, (7–) 15–40 × (2–) 8–18 mm, margins dentate to dentate-serrate; peduncles and pedicels ascending to erect, densely glandular-pubescent. Flowers: calyx lobes ovate to lanceolate, 3–6 × 0.8–1.6 mm, densely glandular-pubescent; corolla white to light lavender or light pink, with reddish violet nectar guides, funnelform, 9–11 mm, glandular-pubescent externally, glandular-pubescent internally, tube 3–4 mm, throat slightly inflated, 3–4 mm diam., rounded abaxially; stamens included, pollen-sacs opposite, navicular to subexplanate, 0.5–0.7 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 9–10 mm, included or exserted, 0.2–0.3 mm diam., tip straight, glabrous; style 7–8 mm. Capsules 4–5 × 2.5–3 mm, glabrous.


Phenology: Flowering Jun–Jul.
Habitat: Igneous soils and crevices, basalt in sagebrush shrublands and pine woodlands.
Elevation: 1200–1800 m.

Discussion

Penstemon sudans is known from northeastern California (Lassen, Modoc, and Placer counties); it has been reported for Washoe County, Nevada. D. D. Keck (1940b) observed that some plants of P. deustus from southeastern Oregon (Curry, Jackson, and Josephine counties) approach P. sudans in glandularity, though he considered these to be one extreme of the morphologic spectrum exhibited by P. deustus.

Selected References

None.

Lower Taxa

None.

"/4+timescorollathroat" is not declared as a valid unit of measurement for this property.

... more about "Penstemon sudans"
acute +  and acuminate +
Craig C. Freeman +
M. E. Jones +
glandular-pubescent +
alternate +, subopposite +, whorled +  and opposite +
glaucescent +
alternate +, opposite +, whorled +, helical +  and subopposite +
cauline +, proximal +  and basal +
rarely +, not glaucous +, glabrate +, pubescent +, hairy +, glandular-lanate +, glandular-pubescent +, puberulent +, glandular +  and glabrous +
not +  and leathery +
tapered +
obovate;ovate or lanceolate +
not +  and leathery +
glandular-pubescent +
ovate;lanceolate +
0.8mm;1.6mm +
short-tubular +  and short-campanulate +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Susanville beardtongue +
personate +, not personate +, bilabiate +  and symmetric +
white +  and light lavender or light pink +
glandular-pubescent +
ventricose +, tubular +  and tubular-funnelform +
ampliate +
0.9 cm9 mm <br />0.009 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
1-5-flowered +
scale-like +  and not needlelike +
Calif. +  and Nev. +
drupe-like +
1200–1800 m. +
curved +  and straight +
1 +  and 4 +
Igneous soils and crevices, basalt in sagebrush shrublands and pine woodlands. +
0 cm0 mm <br />0 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
membranous +
terminal +  and axillary +
subopposite +  and opposite +
persistent +  and deciduous +
glandular-pubescent +
not leathery +
orbiculate;linear;orbiculate;linear;truncate;oblanceolate;obovate;oblong;elliptic;round;lanceolate;ovate +
dentate +  and dentate-serrate +
glutinous +
glandular-ciliolate +, hairy +, puberulent +, glabrescent +, glabrous +  and glandular-pubescent +
lacerate +, toothed +  and entire +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
ascending +  and erect +
glandular-pubescent +
ascending;erect +
glandular-pubescent +
5 +  and 4 +
Flowering Jun–Jul. +
navicular-sigmoid +
1/5 +  and 2/3 +
short-petiolate +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (4 cm40 mm <br />0.04 m <br />) +
ovate +  and lanceolate +
0.8 cm8 mm <br />0.008 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
1.4 cm14 mm <br />0.014 m <br /> (6 cm60 mm <br />0.06 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Contr. W. Bot. +
minute +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
black +, gray +, brown +  and tan +
angled +  and slightly rounded +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
exserted +  and included +
0.9 cm9 mm <br />0.009 m <br /> (?) +  and 1 cm10 mm <br />0.01 m <br /> (?) +
subalternate +, alternate +, whorled +  and opposite +
glutinous +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
ascending;erect +
glaucous +  and glandular-pubescent +
18 cm180 mm <br />0.18 m <br /> (70 cm700 mm <br />0.7 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
papillate +
denticulate +
Penstemon deustus subsp. sudans +
Penstemon sudans +
Penstemon sect. Penstemon +
species +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
rounded +  and inflated +
not constricted +
continuous +  and interrupted +
cylindric +
13 cm130 mm <br />0.13 m <br /> (35 cm350 mm <br />0.35 m <br />) +
straight +
gibbous +  and not spurred +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +