Penstemon xylus

A. Nelson

Bot. Gaz. 34: 31. 1902.

Common names: Tushar Range beardtongue
EndemicConservation concern
Basionym: Penstemon caespitosus var. suffruticosus A. Gray in A. Gray et al. Syn. Fl. N. Amer. 2(1): 270. 1878
Treatment appears in FNA Volume 17. Treatment on page 107. Mentioned on page 99, 103, 108.

Stems prostrate, spreading, or ascending, 6–12 cm, retrorsely hairy, hairs pointed, sometimes also appressed, white, scalelike. Leaves not leathery, glabrous or sparsely hairy, hairs appressed, white, scalelike abaxially, ± densely hairy, hairs appressed, white, scalelike adaxially; cauline 1–5 pairs, petiolate, 6–16 × 2–4.5 mm, blade obovate to spatulate, base tapered, apex rounded to mucronate or acute. Thyrses continuous, secund, 2–12 cm, axis retrorsely hairy, hairs appressed or spreading, white, scalelike, sometimes also glandular-pubescent distally, verticillasters 3–12, cymes 1–3 (or 4) -flowered, 1 per node; proximal bracts spatulate to oblanceolate, 8–17 × 1.4–4 mm; peduncles and pedicels spreading to ascending, glandular-pubescent, hairs appressed, white, scalelike. Flowers: calyx lobes lanceolate to oblong, 3–5 × 0.7–1 mm, herbaceous-margined, glandular-pubescent and retrorsely hairy, hairs appressed, white, scalelike; corolla blue to lavender, or violet, lined internally abaxially with reddish violet nectar guides, ampliate, 12–18 mm, yellow-lanate internally abaxially, tube 5–6 mm, throat gradually inflated, 3.5–5 mm diam., 2-ridged abaxially; stamens reaching orifice or longer pair exserted, filaments glabrous, pollen-sacs opposite, navicular to subexplanate, 0.6–0.8 mm, sutures papillate; staminode 7–8 mm, included, flattened distally, 0.2–0.3 mm diam., tips straight to recurved, distal 6–7 mm pubescent, hairs yellow or orange, to 0.7 mm; style 11–14 mm. Capsules 3–5 × 3–4 mm.


Phenology: Flowering Jun–Aug.
Habitat: Sandy or gravelly slopes, sagebrush shrublands, aspen woodlands, pine woodlands.
Elevation: 2100–3100 m.

Discussion

Penstemon xylus is known from south-central Utah. Populations are documented in Beaver, Garfield, Iron, Piute, and Sevier counties. The placement of P. xylus remains unclear. D. D. Keck (1937) treated it as P. caespitosus subsp. suffruticosus. N. H. Holmgren (1979b) believed it to be more closely related to P. thompsoniae, which also has vestiture of appressed, white, scalelike hairs.

The correct name for Penstemon xylus at the rank of species, which has been a source of confusion, was discussed by C. C. Freeman (2017).

Selected References

None.

Lower Taxa

None.

"/4+timescorollathroat" is not declared as a valid unit of measurement for this property.

... more about "Penstemon xylus"
rounded +  and mucronate or acute +
Craig C. Freeman +
A. Nelson +
alternate +, opposite +, whorled +, helical +  and subopposite +
tapered +
Penstemon caespitosus var. suffruticosus +
obovate;spatulate +
not +  and leathery +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
herbaceous-margined +
hairy +  and glandular-pubescent +
lanceolate;oblong +
short-tubular +  and short-campanulate +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
Tushar Range beardtongue +
blue +  and lavender or violet +
yellow-lanate +
salverform +  and tubular-funnelform +
ampliate +
1.2 cm12 mm <br />0.012 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
1-3-flowered +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
scale-like +  and not needlelike +
drupe-like +
2100–3100 m. +
curved +  and straight +
1 +  and 4 +
Sandy or gravelly slopes, sagebrush shrublands, aspen woodlands, pine woodlands. +
orange +, yellow +  and white +
spreading +  and appressed +
glandular-pubescent +  and hairy +
scale-like +  and pointed +
0 cm0 mm <br />0 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
membranous +
terminal +  and axillary +
persistent +  and deciduous +
cauline +, proximal +  and basal +
hairy +  and glabrous +
not leathery +
orbiculate;linear;orbiculate;linear;truncate;oblanceolate;obovate;oblong;elliptic;round;lanceolate;ovate +
erose +  and entire +
glabrate +, hairy +  and glandular-pubescent +
toothed +  and entire +
herbaceous +  and scarious +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
spreading +  and ascending +
glandular-pubescent +
spreading;ascending +
glandular-pubescent +
5 +  and 4 +
Flowering Jun–Aug. +
navicular-sigmoid +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
spatulate +  and oblanceolate +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
minute +
dark-brown +  and black +
angled +  and rounded +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.21 cm2.1 mm <br />0.0021 m <br />) +
Endemic +  and Conservation concern +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (?) +  and 0.8 cm8 mm <br />0.008 m <br /> (?) +
subalternate +, alternate +, whorled +  and opposite +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
ascending;spreading;ascending;spreading;prostrate +
6 cm60 mm <br />0.06 m <br /> (12 cm120 mm <br />0.12 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
papillate +
denticulate +
Penstemon xylus +
Penstemon sect. Caespitosi +
species +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
2-ridged +  and inflated +
not constricted +
secund +  and continuous +
cylindric +
2 cm20 mm <br />0.02 m <br /> (12 cm120 mm <br />0.12 m <br />) +
straight +
gibbous +  and not spurred +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
subshrub +  and cespitose +