Pentagramma triangularis subsp. triangularis

(Kaulfuss) Yatskievych
Illustrated
Treatment appears in FNA Volume 2.

Petiole glabrous, not viscid-glandular. Blade thin and herbaceous to somewhat thick and leathery, not viscid-glandular, abaxially densely pale to bright-yellow, adaxially glabrous. Distal pinnae mostly regularly lobed. Proximal basiscopic lobes of basal pinnae pinnatifid, often deeply so. 2n = 60, ca. 90, 120.


Habitat: Chaparral, pine and oak woodlands
Elevation: 50–1800 m

Distribution

V2 286-distribution-map.gif

B.C., Calif., Idaho, Oreg., Wash., Mexico in Baja California

Discussion

We here restrict Pentagramma triangularis subsp. triangularis to plants with yellow farina and glabrous adaxial leaf surfaces occurring throughout a large region in westernmost North America. This subspecies comprises a complex of morphological, cytological, and phytochemical variants, at least some of which may deserve formal taxonomic recognition, following more detailed studies. Plants with yellow farina reported from Arizona, Nevada, and Utah may represent tetraploid hybrids between P. triangularis subsp. triangularis and P. triangularis subsp. maxonii and are not mapped herein.

Selected References

None.

Lower Taxa

None.

"thick" is not a number.

120 +, 90 +  and 60 +
George Yatskievych +  and Michael D. Windham +
(Kaulfuss) Yatskievych +
Gymnogramma triangularis +
abaxially densely pale;bright-yellow +
pinnatifid +, 1-2-pinnate-pinnatifid +  and triangular-pentagonal +
B.C. +, Calif. +, Idaho +, Oreg. +, Wash. +  and Mexico in Baja California +
50–1800 m +
yellow adaxially bright green +  and yellowish green +
glabrous +  and glandular +
obcordate;reniform +
Chaparral, pine and oak woodlands +
monomorphic +
recurved +  and reflexed +
noncircinate +  and circinate +
5 cm50 mm <br />0.05 m <br /> (40 cm400 mm <br />0.4 m <br />) +
not recurved +
chestnut-brown;dark-brown +
nearly +  and rounded +
Amer. Fern J. +
bicolored with black +
persistent +
Illustrated +
tan +  and brown +
trigonal +, globose +  and globose-tetrahedral +
ascending +, erect +  and short-creeping +
Pityrogramma triangularis +
Pentagramma triangularis subsp. triangularis +
Pentagramma triangularis +
subspecies +
crescent--shaped +  and roundish +