Physaria arizonica

(S. Watson) O’Kane & Al-Shehbaz

Novon 12: 321. 2002.

Common names: Arizona bladderpod
Basionym: Lesquerella arizonica S. Watson Proc. Amer. Acad. Arts 23: 254. 1888
Synonyms: Lesquerella arizonica var. nudicaulis Payson Physaria arizonica var. andrusensis N. D. Atwood, S. L. Welsh & L.C. Higgins
Treatment appears in FNA Volume 7. Treatment on page 627. Mentioned on page 621.

Perennials; caudex branched, (cespitose); densely (silvery gray) pubescent, trichomes (sessile or subsessile), (4-), 6-rayed, or 8-rayed, rays fused at base, furcate or bifurcate, (slightly umbonate, tuberculate throughout). Stems simple or few to several from base, erect, (unbranched, slender), 0.2–1 (–1.5) dm. Basal leaves (densely tufted, not rosulate, reflexed in age); blade obovate to oblanceolate, 0.7–2 (–3) cm, margins usually entire, sometimes repand or shallowly dentate, (apex acute). Cauline leaves similar to basal, becoming narrower distally, somewhat reflexed, (distal) blade linear or narrowly oblanceolate, 0.5–2.5 (–5.5) cm. Racemes dense, often subcorymbiform. Fruiting pedicels (erect or divaricate-spreading, straight or slightly curved), (3–) 5–10 (–15) mm. Flowers: sepals (green or greenish yellow), ovate or broadly ovate, 3.5–6.5 mm, (lateral pair subsaccate, cucullate, median pair thickened, slightly cucullate apically); petals (spreading), oblanceolate to obovate, 5.5–8 (–10) mm, (claw erect). Fruits (sessile or substipitate), suborbicular to ovoid or ellipsoid, slightly inflated, 4–7 mm; valves pubescent outside, trichomes substipitate, spreading, sometimes sparsely pubescent inside, trichomes sessile, smooth; ovules 4–10 (–16) per ovary; style (0.5–) 1–2 (–4) mm (shorter than fruit). Seeds flattened. 2n = 10.


Phenology: Flowering Apr–Jun.
Habitat: Sandy and gravelly soils, limey knolls or limestone chip, often in open stands of sagebrush-pinyon, pinyon-juniper, Gambel oak and sometimes ponderosa pine
Elevation: 1000-2200 m

Discussion

The circumscription of Physaria arizonica here is quite broad and includes plants that have densely tufted basal leaves and relatively few or no cauline leaves; plants that are loosely tufted and have several cauline leaves; and plants that have a strongly branched caudex, leafy stems, and sterile shoots (var. andrusensis). Additional study is needed to understand the pattern of variation in this complex species; all of the characters given above vary considerably.

Selected References

None.

Lower Taxa

None.

"narrower" is not a number."not" is not a number. "elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Physaria arizonica"
ovate +  and narrowly oblong +
Steve L. O’Kane Jr. +
(S. Watson) O’Kane & Al-Shehbaz +
Lesquerella arizonica +
appendaged +  and unappendaged +
2.5 cm25 mm <br />0.025 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
repand +  and dentate +
oblanceolate;linear;obovate;oblanceolate +
much smaller +  and reduced +
0.5 cm5 mm <br />0.005 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
urceolate +, campanulate +  and tubular +
sessile +  and petiolate +
well-developed +
distinct +
not +  and differentiated +
Arizona bladderpod +
emarginate +  and entire +
Ariz. +  and Utah. +
1000-2200 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
oblanceolate +  and obovate +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
2-valved +  and capsular +
inflated +, suborbicular +  and ovoid or ellipsoid +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
latiseptate +, stipitate +, , +, subsessile +  and sessile +
angustiseptate +, terete +, not +, inflated +, not torulose +, obovoid +, ovoid +, ovate +, obpyriform +, oblong +, obdeltate +, obcordate +, lanceolate +, elliptic +, ellipsoid +, suborbicular +, orbicular +, subglobose +  and globose +
latiseptate +  and unsegmented +
angustiseptate +  and terete +
Sandy and gravelly soils, limey knolls or limestone chip, often in open stands of sagebrush-pinyon, pinyon-juniper, Gambel oak and sometimes ponderosa pine +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
dentate;repand +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
purple +, white +, orange +  and yellow +
rudimentary +
obdeltate +, oblanceolate +, ovate +, obovate +  and spatulate +
Flowering Apr–Jun. +
trinucleate +  and 3(-11)-colpate +
subcorymbiform +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
ascending +, spreading +, horizontal +  and erect +
slender +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
bifurcate +  and furcate +
rounded +  and narrowly oblong +
not +  and mucilaginous +
margined +  and winged +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
flattened +
persistent +  and caducous +
distinct +
spreading +  and erect +
not saccate +, subsaccate +, deltate +, ovate +, oblong +, elliptic +, lanceolate +  and linear +
perforate +  and complete +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
few to several +  and simple +
procumbent +  and decumbent +
prostrate +  and decumbent +
2-lobed +  and entire +
subsessile +, sessile +  and petiolate +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
persistent +
distinct +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Lesquerella arizonica var. nudicaulis +  and Physaria arizonica var. andrusensis +
Physaria arizonica +
Physaria +
species +
sessile +  and substipitate +
stellate-scalelike +
coiled +, 1-7-veined +  and veined +
anastomosing +
perennial +, biennial +  and annual +
aquatics +  and terrestrial +
glabrous +  and pubescent +
15 +, 12 +, 10 +, 9 +, 8 +, 7 +, 6 +, 5 +  and 4 +