Physaria didymocarpa

(Hooker) A. Gray

Gen. Amer. Bor. 1: 162. 1848.

Common names: Common twinpod
Endemic
Basionym: Vesicaria didymocarpa Hooker Fl. Bor.-Amer. 1: 49, plate 16. 1830
Synonyms: Coulterina didymocarpa (Hooker) Kuntze
Treatment appears in FNA Volume 7. Treatment on page 633. Mentioned on page 618, 644, 661.

Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). Stems several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm. Basal leaves (forming a strong rosette; long-petiolate); blade obovate, 1.5–4 (–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery). Cauline leaves: blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute). Racemes congested, (elongated in fruit, greatly exceeding leaves). Fruiting pedicels (spreading, straight or slightly curved), 8–12 mm. Flowers: sepals lanceolate to oblong, 6–8 mm, (often keeled); petals spatulate, 10–12 mm. Fruits (erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy); replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse; ovules (4–) 8 per ovary; style 7–9 mm. Seeds flattened.

Distribution

V7 1039-distribution-map.gif

Alta., B.C., Idaho, Mont., Wash., Wyo.

Discussion

Subspecies 3 (3 in the flora).

The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa.

Selected References

None.

Key

1 Trichomes spreading throughout; basal leaf blade margins dentate. Physaria didymocarpa subsp. lanata
1 Trichomes appressed (except fruits in subsp. didymocarpa); basal leaf blade margins repand to dentate, ± lyrate or, rarely, entire > 2
2 Fruits moderately inflated, trichomes spreading. Physaria didymocarpa subsp. didymocarpa
2 Fruits highly inflated (papery), trichomes appressed. Physaria didymocarpa subsp. lyrata

No values specified."not" is not a number. "elongated" is not a number."thick" is not a number.

... more about "Physaria didymocarpa"
ovate +  and narrowly oblong +
Steve L. O’Kane Jr. +
(Hooker) A. Gray +
Vesicaria didymocarpa +
appendaged +  and unappendaged +
4 cm40 mm <br />0.04 m <br /> (8 cm80 mm <br />0.08 m <br />) +
repand +  and dentate +
oblanceolate;obovate +
much smaller +  and reduced +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
urceolate +, campanulate +  and tubular +
sessile +  and petiolate +
well-developed +
distinct +
not +  and differentiated +
Common twinpod +
emarginate +  and entire +
Alta. +, B.C. +, Idaho +, Mont. +, Wash. +  and Wyo. +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
2-valved +  and capsular +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
inflated +
latiseptate +, stipitate +, , +, subsessile +  and sessile +
angustiseptate +, terete +, not +, inflated +, not torulose +, obovoid +, ovoid +, ovate +, obpyriform +, oblong +, obdeltate +, obcordate +, lanceolate +, elliptic +, ellipsoid +, suborbicular +, orbicular +, subglobose +  and globose +
latiseptate +  and unsegmented +
angustiseptate +  and terete +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
with occasional tooth +, entire +  and dentate +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
purple +, white +, orange +  and yellow +
rudimentary +
spatulate +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
trinucleate +  and 3(-11)-colpate +
Gen. Amer. Bor. +
ascending +, spreading +, horizontal +  and erect +
slender +
0.8 cm8 mm <br />0.008 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
furcate +
obovate +  and broadly oblong +
not constricted +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
not +  and mucilaginous +
margined +  and winged +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
flattened +
persistent +  and caducous +
distinct +
spreading +  and erect +
lanceolate +  and oblong +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
perforate +  and complete +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
spreading +  and erect +
1 cm10 mm <br />0.01 m <br /> (?) +
2-lobed +  and entire +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.7 cm7 mm <br />0.007 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
Coulterina didymocarpa +
Physaria didymocarpa +
Physaria +
species +
stellate-scalelike +
coiled +, 1-7-veined +  and veined +
anastomosing +
perennial +, biennial +  and annual +
aquatics +  and terrestrial +
glabrous +  and pubescent +
15 +, 12 +, 10 +, 9 +, 8 +, 7 +, 6 +, 5 +  and 4 +