Physaria douglasii

(S. Watson) O’Kane & Al-Shehbaz

Novon 12: 322. 2002.

Common names: Douglas’s bladderpod
Basionym: Lesquerella douglasii S. Watson Proc. Amer. Acad. Arts 23: 255. 1888
Treatment appears in FNA Volume 7. Treatment on page 634. Mentioned on page 623.

Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6 (–10) -rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). Stems simple from base, erect, (usually unbranched), to 4.5 dm. Basal leaves: blade suborbicular to elliptic, 2–9.5 (–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. Cauline leaves similar to basal, blade narrowly linear or, sometimes, orbicular. Racemes loose (lax). Fruiting pedicels (recurved, straight, curved, or sigmoid), 6–20 mm. Flowers: sepals elliptic or ovate, (2–) 3.5–7.5 mm, (cucullate); petals 6–11 mm. Fruits obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4 (–8) per ovary; style (1.6–) 3–6 mm. Seeds flattened. 2n = 10, 30.

Distribution

V7 1044-distribution-map.gif

B.C., Idaho, Mont., Oreg., Wash.

Discussion

Subspecies 2 (2 in the flora).

Selected References

None.

Key

1 Fruit valves: trichomes sessile; cauline leaves loosely arranged, blades narrowly linear; British Columbia, Idaho, Montana, Oregon, Washington. Physaria douglasii subsp. douglasii
1 Fruit valves: trichomes stalked; cauline leaves imbricate, blades sometimes orbicular; White Bluffs adjacent to Columbia River of Washington. Physaria douglasii subsp. tuplashensis

"not" is not a number. "elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property.

... more about "Physaria douglasii"
ovate +  and narrowly oblong +
Steve L. O’Kane Jr. +
(S. Watson) O’Kane & Al-Shehbaz +
Lesquerella douglasii +
appendaged +  and unappendaged +
9.5 cm95 mm <br />0.095 m <br /> (11.5 cm115 mm <br />0.115 m <br />) +
repand +  and dentate +
orbicular;suborbicular;elliptic +
much smaller +  and reduced +
2 cm20 mm <br />0.02 m <br /> (9.5 cm95 mm <br />0.095 m <br />) +
urceolate +, campanulate +  and tubular +
sessile +  and petiolate +
well-developed +
distinct +
not +  and differentiated +
Douglas’s bladderpod +
emarginate +  and entire +
B.C. +, Idaho +, Mont. +, Oreg. +  and Wash. +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
2-valved +  and capsular +
not inflated +, obovoid +  and subglobose +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
latiseptate +, stipitate +, , +, subsessile +  and sessile +
angustiseptate +, terete +, not +, inflated +, not torulose +, obovoid +, ovoid +, ovate +, obpyriform +, oblong +, obdeltate +, obcordate +, lanceolate +, elliptic +, ellipsoid +, suborbicular +, orbicular +, subglobose +  and globose +
latiseptate +  and unsegmented +
angustiseptate +  and terete +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
lyrate-pinnatifid +, dentate +  and entire +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
purple +, white +, orange +  and yellow +
rudimentary +
obdeltate +, oblanceolate +, ovate +, obovate +  and spatulate +
0.6 cm6 mm <br />0.006 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
trinucleate +  and 3(-11)-colpate +
ascending +, spreading +, horizontal +  and erect +
slender +
0.6 cm6 mm <br />0.006 m <br /> (2 cm20 mm <br />0.02 m <br />) +
bifurcate +  and furcate +
rounded +  and narrowly oblong +
not +  and mucilaginous +
margined +  and winged +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
flattened +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
persistent +  and caducous +
distinct +
spreading +  and erect +
ovate +  and elliptic +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
perforate +  and complete +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
procumbent +  and decumbent +
prostrate +  and decumbent +
2-lobed +  and entire +
subsessile +, sessile +  and petiolate +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
persistent +
distinct +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Coulterina +  and Lesquerella +
Physaria douglasii +
Physaria +
species +
stalked +  and sessile +
stellate-scalelike +
coiled +, 1-7-veined +  and veined +
glabrous +  and pubescent +
anastomosing +
perennial +, biennial +  and annual +
aquatics +  and terrestrial +
glabrous +  and pubescent +
15 +, 12 +, 10 +, 9 +, 8 +, 7 +, 6 +, 5 +  and 4 +