Physaria hitchcockii

(Munz) O’Kane & Al-Shehbaz

Novon 12: 324. 2002.

Common names: Hitchcock’s bladderpod
Basionym: Lesquerella hitchcockii Munz Bull. Torrey Bot. Club 56: 163. 1929
Treatment appears in FNA Volume 7. Treatment on page 643. Mentioned on page 619, 664.

Perennials; (forming loose mats); caudex (buried), branched; densely pubescent, trichomes (short-stalked), 4–6-rayed, rays distinct, bifurcate, (rough-tuberculate). Stems few to several from base, prostrate to erect or spreading, 0.05–0.5 (–1.2) dm. Basal leaves: (petiole and blade differentiated or not); blade spatulate to elliptic or linear or narrowly oblanceolate, 0.5–1.5 (–2.5) cm, margins entire. Cauline leaves similar to basal, smaller. Racemes dense. Fruiting pedicels (ascending, straight or slightly curved), 2–6 mm. Flowers: sepals narrowly lanceolate to lanceolate, 2.8–6 mm; petals (pale to deep yellow), narrowly lanceolate to oblanceolate, 5–9 mm, (claw undifferentiated from blade). Fruits (sessile or substipitate), globose or subglobose to obovoid, not or slightly inflated, 3–6 mm, (firm, apex acute); valves (reddish in age, not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8 per ovary; style 1.7–6 mm. Seeds flattened.

Discussion

Subspecies 3 (3 in the flora).

The taxonomic treatment of Physaria hitchcockii has varied widely over the years. Molecular study (pers. obs.) has shown no direct relationship to P. tumulosa; morphologically, though, P. navajoensis and P. tumulosa appear closely related. Infraspecific taxonomy is based on the presence of a discernable petiole and whether or not the caudex is elastically elongated. The subspecies recognized here are usually geographically coherent, except that collections from the Table Cliff Plateau are more similar to subsp. hitchcockii, disjunct in Nevada, than they are to the very nearly sympatric subsp. rubicundula.

Selected References

None.

Key

1 Caudices elongated and elastic; basal leaves: petiole not differentiated from blade, blades linear-oblanceolate; Aquarius, Markagunt, and Paunsaugunt plateaus, Utah (limited to the pink member of the limestone Wasatch (Claron) Formation). Physaria hitchcockii subsp. rubicundula
1 Caudices elongated or not, elastic or not; basal leaves: petiole differentiated (sometimes weakly) from blade, blades oblanceolate to obovate; Nevada, Utah > 2
2 Plants forming tufts; caudices not elongated, not elastic; fruits 2.6-3.8 mm wide; Table Cliff Plateau, Utah (limited to the white member of the limestone Wasatch (Claron) Formation) and limestones of the Sheep Range and Spring Mountains, Nevada. Physaria hitchcockii subsp. hitchcockii
2 Plants forming soft mats; caudices elongated, elastic (creeping); fruits 1.7-3 mm wide; Grant, Quinn Canyon, and Schell Creek ranges, Nevada. Physaria hitchcockii subsp. confluens

"not" is not a number. "elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Physaria hitchcockii"
ovate +  and narrowly oblong +
Steve L. O’Kane Jr. +
(Munz) O’Kane & Al-Shehbaz +
Lesquerella hitchcockii +
appendaged +  and unappendaged +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
repand +  and dentate +
linear or narrowly oblanceolate;spatulate to elliptic +
much smaller +  and reduced +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
urceolate +, campanulate +  and tubular +
sessile +  and petiolate +
well-developed +
distinct +
not +  and differentiated +
Hitchcock’s bladderpod +
emarginate +  and entire +
Nev. +  and Utah +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
narrowly lanceolate +  and oblanceolate +
0.5 cm5 mm <br />0.005 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
2-valved +  and capsular +
inflated +, subglobose +  and obovoid +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
latiseptate +, stipitate +, , +, subsessile +  and sessile +
angustiseptate +, terete +, not +, inflated +, not torulose +, obovoid +, ovoid +, ovate +, obpyriform +, oblong +, obdeltate +, obcordate +, lanceolate +, elliptic +, ellipsoid +, suborbicular +, orbicular +, subglobose +  and globose +
latiseptate +  and unsegmented +
angustiseptate +  and terete +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
sinuate;dentate;sinuate;dentate +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
purple +, white +, orange +  and yellow +
rudimentary +
obdeltate +, oblanceolate +, ovate +, obovate +  and spatulate +
trinucleate +  and 3(-11)-colpate +
ascending +, spreading +, horizontal +  and erect +
slender +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
distinct +
bifurcate +
rounded +  and narrowly oblong +
not +  and mucilaginous +
margined +  and winged +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
flattened +
persistent +  and caducous +
distinct +
spreading +  and erect +
narrowly lanceolate +  and lanceolate +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
perforate +  and complete +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
prostrate +  and decumbent +
prostrate +  and erect or spreading +
few +  and several +
2-lobed +  and entire +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.17 cm1.7 mm <br />0.0017 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Coulterina +  and Lesquerella +
Physaria hitchcockii +
Physaria +
species +
simple +, stalked +, subsessile +  and sessile +
stellate-scalelike +
coiled +, 1-7-veined +  and veined +
anastomosing +
perennial +, biennial +  and annual +
aquatics +  and terrestrial +
glabrous +  and pubescent +
15 +, 12 +, 10 +, 9 +, 8 +, 7 +, 6 +, 5 +  and 4 +