Physaria integrifolia

(Rollins) Lichvar

Madroño 31: 203. 1984.

Common names: Snake River or creeping twinpod
Basionym: Physaria didymocarpa var. integrifolia Rollins Rhodora 41: 407. 1939
Synonyms: Physaria integrifolia var. monticola Lichvar
Treatment appears in FNA Volume 7. Treatment on page 644. Mentioned on page 618.

Perennials; caudex usually branched, (rhizomelike, cespitose); densely (silvery) pubescent, trichomes (often stalked, appressed), several-rayed, rays furcate or bifurcate, (umbonate, strongly tuberculate throughout). Stems several from base, ± erect, exceeding basal rosette by ± 0.5 dm. Basal leaves (forming a strong rosette; long-petiolate); blade oblanceolate to ovate or orbicular, (1.5–) 2–4 (–8) cm, (base usually abruptly tapering to petiole), margins entire. Cauline leaves: blade oblanceolate, 1–2 cm, margin entire, (apex acute). Racemes congested, (greatly exceeding leaves). Fruiting pedicels (spreading, straight or slightly curved), 7–11 mm. Flowers: sepals often keeled, 6–8 mm; petals spatulate, 8–10 mm. Fruits didymous, highly inflated, 8–22 × 10–25 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), densely pubescent, trichomes appressed; replum linear to oblong, as wide as or wider than fruit; ovules 8 per ovary; style 7–9 mm. Seeds flattened. 2n = 16.


Phenology: Flowering (May-)Jun–Jul(-Aug).
Habitat: Calcareous hills and slopes, shale-limestone cliffs, bare steep slopes, red clay banks, shale
Elevation: 1900-2700 m

Discussion

Physaria integrifolia has traditionally been recognized as a variety of P. didymocarpa, but it is morphologically and ecologically quite distinctive. Variety monticola (no combination has been made at subspecific rank) is not recognized here; it is considered another example, in the genus, of caudices elongating in response to shifting substrates.

Selected References

None.

Lower Taxa

None.

No values specified."not" is not a number. "elongated" is not a number."thick" is not a number.

... more about "Physaria integrifolia"
ovate +  and narrowly oblong +
Steve L. O’Kane Jr. +
(Rollins) Lichvar +
Physaria didymocarpa var. integrifolia +
appendaged +  and unappendaged +
4 cm40 mm <br />0.04 m <br /> (8 cm80 mm <br />0.08 m <br />) +
repand +  and dentate +
oblanceolate;oblanceolate;ovate or orbicular +
much smaller +  and reduced +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
urceolate +, campanulate +  and tubular +
sessile +  and petiolate +
well-developed +
distinct +
not +  and differentiated +
Snake River or creeping twinpod +
emarginate +  and entire +
Idaho +, Mont. +  and Wyo. +
1900-2700 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
2-valved +  and capsular +
0.8 cm8 mm <br />0.008 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
inflated +
latiseptate +, stipitate +, , +, subsessile +  and sessile +
angustiseptate +, terete +, not +, inflated +, not torulose +, obovoid +, ovoid +, ovate +, obpyriform +, oblong +, obdeltate +, obcordate +, lanceolate +, elliptic +, ellipsoid +, suborbicular +, orbicular +, subglobose +  and globose +
latiseptate +  and unsegmented +
angustiseptate +  and terete +
Calcareous hills and slopes, shale-limestone cliffs, bare steep slopes, red clay banks, shale +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
sinuate;dentate;sinuate;dentate +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
purple +, white +, orange +  and yellow +
rudimentary +
spatulate +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Flowering (May-)Jun–Jul(-Aug). +
trinucleate +  and 3(-11)-colpate +
ascending +, spreading +, horizontal +  and erect +
slender +
0.7 cm7 mm <br />0.007 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
bifurcate +  and furcate +
linear +  and oblong +
not +  and mucilaginous +
margined +  and winged +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
flattened +
persistent +  and caducous +
distinct +
spreading +  and erect +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
perforate +  and complete +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
prostrate +  and decumbent +
2-lobed +  and entire +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.7 cm7 mm <br />0.007 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
Physaria integrifolia var. monticola +
Physaria integrifolia +
Physaria +
species +
stellate-scalelike +
coiled +, 1-7-veined +  and veined +
anastomosing +
perennial +, biennial +  and annual +
aquatics +  and terrestrial +
glabrous +  and pubescent +
15 +, 12 +, 10 +, 9 +, 8 +, 7 +, 6 +, 5 +  and 4 +