Physaria intermedia

(S. Watson) O’Kane & Al-Shehbaz

Novon 12: 324. 2002.

Common names: Mid-bladderpod
Basionym: Lesquerella alpina var. intermedia S. Watson Proc. Amer. Acad. Arts 23: 251. 1888
Synonyms: Lesquerella intermedia (S. Watson) A. Heller
Treatment appears in FNA Volume 7. Treatment on page 644. Mentioned on page 621, 645, 651.

Perennials; caudex (buried), branched, (thickened with persistent leaf-bases, cespitose); densely pubescent (usually grayish-green), trichomes (sessile or short stalked, spreading), several-rayed, rays furcate or bifurcate, slightly fused at base, (tuberculate or finely tuberculate). Stems several from base, erect to decumbent, (unbranched, stout, densely leafy sterile shoots sometimes present), (0.5–) 4–2.5 dm. Basal leaves (clustered at stem base); blade linear to linear-oblanceolate, 2–5 cm, margins entire, usually involute, sometimes flattened, (apex obtuse to subacute). Cauline leaves: blade linear-oblanceolate to linear, 1–3.5 (–4.5) cm, margins entire, usually involute. Racemes compact, (often nearly subumbellate). Fruiting pedicels (often expanded distally, ascending or recurved, usually straight or slightly curved, rarely nearly sigmoid), 4–15 mm, (stout). Flowers: sepals (yellowish or greenish yellow), ovate or oblong, 4.5–7.5 (–9) mm, (lateral pair sometimes cucullate, median pair tapering at both ends, thickened apically, cucullate); petals spatulate or oblong, 6.5–10.5 (–15) mm, (base sometimes widened, apex rounded or retuse). Fruits (sessile or substipitate), subglobose to slightly ovoid, usually inflated, rarely compressed or obcompressed, 4–6 (–10) mm, (apex acute, slightly flattened); valves sparsely pubescent, trichomes appressed; ovules (8–) 12–16 (–20) per ovary; style (2–) 3–4.5 (–5.5) mm. Seeds flattened. 2n = 18, 20, 36.


Phenology: Flowering Apr–Aug.
Habitat: Dry sandy, gravelly, or rocky soil, claylike hillsides, open chiprock, dry stream beds, gravel bars, open knolls, open pinyon-juniper woods, open stands of sagebrush, Gambel oak or ponderosa pine communities, calcareous substrates
Elevation: 1600-2400 m

Discussion

N. H. Holmgren (2005b) pointed out that the lectotype and other material from New Mexico, where Physaria intermedia is very infrequent, is quite similar to P. parvula from northern Colorado and northeastern Utah; it is also quite similar to, but less robust than, P. pulvinata from southwestern Colorado. The material from Arizona, northwestern New Mexico, and Utah may represent an unnamed taxon; further study is needed.

Selected References

None.

Lower Taxa

None.

"not" is not a number. "elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Physaria intermedia"
36 +, 20 +  and 18 +
ovate +  and narrowly oblong +
Steve L. O’Kane Jr. +
(S. Watson) O’Kane & Al-Shehbaz +
Lesquerella alpina var. intermedia +
appendaged +  and unappendaged +
3.5 cm35 mm <br />0.035 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
repand +  and dentate +
linear-oblanceolate;linear +
much smaller +  and reduced +
1 cm10 mm <br />0.01 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
urceolate +, campanulate +  and tubular +
sessile +  and petiolate +
well-developed +
distinct +
not +  and differentiated +
Mid-bladderpod +
emarginate +  and entire +
Ariz. +, N.Mex. +  and Utah. +
1600-2400 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
0.75 cm7.5 mm <br />0.0075 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
oblong +  and ovate +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
2-valved +  and capsular +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
subglobose +  and slightly ovoid usually inflated rarely compressed or obcompressed +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
latiseptate +, stipitate +, , +, subsessile +  and sessile +
angustiseptate +, terete +, not +, inflated +, not torulose +, obovoid +, ovoid +, ovate +, obpyriform +, oblong +, obdeltate +, obcordate +, lanceolate +, elliptic +, ellipsoid +, suborbicular +, orbicular +, subglobose +  and globose +
latiseptate +  and unsegmented +
angustiseptate +  and terete +
Dry sandy, gravelly, or rocky soil, claylike hillsides, open chiprock, dry stream beds, gravel bars, open knolls, open pinyon-juniper woods, open stands of sagebrush, Gambel oak or ponderosa pine communities, calcareous substrates +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
flattened +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
1.05 cm10.5 mm <br />0.0105 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
purple +, white +, orange +  and yellow +
rudimentary +
oblong +  and spatulate +
0.65 cm6.5 mm <br />0.0065 m <br /> (1.05 cm10.5 mm <br />0.0105 m <br />) +
Flowering Apr–Aug. +
trinucleate +  and 3(-11)-colpate +
ascending +, spreading +, horizontal +  and erect +
slender +
0.4 cm4 mm <br />0.004 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
bifurcate +  and furcate +
rounded +  and narrowly oblong +
not +  and mucilaginous +
margined +  and winged +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
flattened +
persistent +  and caducous +
distinct +
spreading +  and erect +
not saccate +, subsaccate +, deltate +, ovate +, oblong +, elliptic +, lanceolate +  and linear +
perforate +  and complete +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
prostrate +  and decumbent +
erect +  and decumbent +
2-lobed +  and entire +
subsessile +, sessile +  and petiolate +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
persistent +
distinct +
0.3 cm3 mm <br />0.003 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
Lesquerella intermedia +
Physaria intermedia +
Physaria +
species +
stellate-scalelike +
coiled +, 1-7-veined +  and veined +
anastomosing +
perennial +, biennial +  and annual +
aquatics +  and terrestrial +
glabrous +  and pubescent +
15 +, 12 +, 10 +, 9 +, 8 +, 7 +, 6 +, 5 +  and 4 +