Physaria ludoviciana

(Nuttall) O’Kane & Al-Shehbaz

Novon 12: 325. 2002.

Common names: Foothill bladderpod
Basionym: Alyssum ludovicianum Nuttall Gen. N. Amer. Pl. 2: 63. 1818,
Synonyms: Lesquerella ludoviciana (Nuttall) S. Watson Vesicaria ludoviciana (Nuttall) de Candolle
Treatment appears in FNA Volume 7. Treatment on page 649. Mentioned on page 620, 650.

Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout). Stems few from base, erect with outer usually decumbent, 1–3.5 (–5) dm. Basal leaves (erect); blade narrowly lanceolate to linear or (outer) oblanceolate, (1–) 2–6 (–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes). Cauline leaves: blade narrowly oblanceolate to linear, (1–) 2–4 (–8) cm, margins flat or involute. Racemes compact, (elongated and loose in fruit, densely-flowered). Fruiting pedicels (usually recurved), (5–) 10–20 (–25) mm. Flowers: sepals oblong to broadly elliptic, 4–7 (–8) mm, (lateral pair subsaccate, median pair cucullate); petals oblanceolate or obovate, (5–) 6.5–9.5 (–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw). Fruits subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–) 4–6 mm; valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile; ovules (4–) 8–12 (–16) per ovary; style 3–4.5 (–6.5) mm. Seeds slightly flattened. 2n = 10, 20, 30.


Phenology: Flowering Apr–Jul(-Aug).
Habitat: Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands
Elevation: 0-1900 m

Distribution

V7 1092-distribution-map.gif

Ariz., Calif., Colo., Ill., Iowa, Kans., Minn., Mont., Nebr., Nev., N.Mex., N.Dak., Okla., S.Dak., Utah, Wis., Wyo.

Discussion

Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana.

Selected References

None.

Lower Taxa

None.

"not" is not a number. "elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Physaria ludoviciana"
30 +, 20 +  and 10 +
ovate +  and narrowly oblong +
Steve L. O’Kane Jr. +
(Nuttall) O’Kane & Al-Shehbaz +
Alyssum ludovicianum +
appendaged +  and unappendaged +
4 cm40 mm <br />0.04 m <br /> (8 cm80 mm <br />0.08 m <br />) +
repand +  and dentate +
narrowly oblanceolate;linear +
much smaller +  and reduced +
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
sessile +  and petiolate +
well-developed +
distinct +
not +  and differentiated +
Foothill bladderpod +
emarginate +  and entire +
Ariz. +, Calif. +, Colo. +, Ill. +, Iowa +, Kans. +, Minn. +, Mont. +, Nebr. +, Nev. +, N.Mex. +, N.Dak. +, Okla. +, S.Dak. +, Utah +, Wis. +  and Wyo. +
0-1900 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
2-valved +  and capsular +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
compressed +, inflated +, obovoid +  and subglobose +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
latiseptate +, stipitate +, , +, subsessile +  and sessile +
angustiseptate +, terete +, not +, inflated +, not torulose +, obovoid +, ovoid +, ovate +, obpyriform +, oblong +, obdeltate +, obcordate +, lanceolate +, elliptic +, ellipsoid +, suborbicular +, orbicular +, subglobose +  and globose +
latiseptate +  and unsegmented +
angustiseptate +  and terete +
Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
dentate +  and entire +
involute;flat +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
0.95 cm9.5 mm <br />0.0095 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
purple +, white +, orange +  and yellow +
rudimentary +
obovate +  and oblanceolate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.95 cm9.5 mm <br />0.0095 m <br />) +
Flowering Apr–Jul(-Aug). +
trinucleate +  and 3(-11)-colpate +
2 cm20 mm <br />0.02 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
ascending +, spreading +, horizontal +  and erect +
slender +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
bifurcate +  and furcate +
rounded +  and narrowly oblong +
not +  and mucilaginous +
margined +  and winged +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
flattened +
0.7 cm7 mm <br />0.007 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
persistent +  and caducous +
distinct +
spreading +  and erect +
oblong +  and broadly elliptic +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
perforate +  and complete +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
prostrate +  and decumbent +
2-lobed +  and entire +
subsessile +, sessile +  and petiolate +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
persistent +
distinct +
0.3 cm3 mm <br />0.003 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
Lesquerella ludoviciana +  and Vesicaria ludoviciana +
Physaria ludoviciana +
Physaria +
species +
stellate-scalelike +
coiled +, 1-7-veined +  and veined +
anastomosing +
perennial +, biennial +  and annual +
aquatics +  and terrestrial +
glabrous +  and pubescent +
15 +, 12 +, 10 +, 9 +, 8 +, 7 +, 6 +, 5 +  and 4 +