Physaria navajoensis

(O’Kane) O’Kane & Al-Shehbaz

Novon 12: 325. 2002.

Common names: Navajo bladderpod
Basionym: Lesquerella navajoensis O’Kane Madroño 46: 88, figs. 1, 2. 1999
Treatment appears in FNA Volume 7. Treatment on page 651. Mentioned on page 619, 643, 664.

Perennials; caudex branched, (woody, pulvinate-cespitose, forming hard, hemispherical mats, basal parts covered with persistent leaf-bases); densely (silvery gray) pubescent, trichomes mostly 5-rayed, rays bifurcate, slightly fused at base, (umbonate, strongly tuberculate except nearly smooth over umbo). Stems several from base (crowded), erect, not exceeding leaves. Basal leaves: usually absent. Cauline leaves: (petiole not differentiated from blade); blade linear-oblanceolate, 3–8 (–13) mm, margins entire. Racemes (secund), dense, corymbose, (few-flowered, not or barely exceeding leaves). Fruiting pedicels (ascending to divaricate-ascending, straight), 3.5–6 mm. Flowers: sepals (yellow-green), linear to narrowly triangular, 3.7–4.8 mm, (lateral pair subsaccate); petals (deep yellow, slightly orange in center), spatulate, 5.2–6.5 mm, (claw joined at right angle). Fruits (becoming reddish or copper-colored in age), ovate, often slightly compressed (at margins apically), 3–5 mm, (apex acute); valves (not retaining seeds after dehiscence), glabrous; (septum perforate or not); ovules 4–8 per ovary; style 1.8–3 mm. Seeds plump or slightly flattened, (strongly mucilaginous).


Phenology: Flowering May-early Jun.
Habitat: Pinyon-juniper communities on nearly barren outcrops of Todilto Limestone
Elevation: 2200-2400 m

Discussion

Of conservation concern.

Physaria navajoensis is morphologically similar to 85. P. tumulosa of southern Utah, differing subtly. Physaria navajoensis has petals slightly orange at the junction of blade and claw, a sharp bend at that junction giving the flower a flat-topped appearance, and strongly mucilaginous seeds. Physaria tumulosa has pure yellow petals that gently flex at the junction of blade and claw, and seeds that are not mucilaginous. Molecular data (pers. obs.) show that these two species are not directly related. A population of plants on Deer Spring Point, Kane County, Utah, appears to be this species, but molecular data indicate that it is probably a hybrid between P. tumulosa and, most likely, P. intermedia.

Selected References

None.

Lower Taxa

None.

"not" is not a number. "elongated" is not a number."thick" is not a number.

... more about "Physaria navajoensis"
ovate +  and narrowly oblong +
Steve L. O’Kane Jr. +
(O’Kane) O’Kane & Al-Shehbaz +
Lesquerella navajoensis +
appendaged +  and unappendaged +
0.8 cm8 mm <br />0.008 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
repand +  and dentate +
linear-oblanceolate +
much smaller +  and reduced +
0.3 cm3 mm <br />0.003 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
urceolate +, campanulate +  and tubular +
sessile +  and petiolate +
well-developed +
distinct +
not +  and differentiated +
Navajo bladderpod +
emarginate +  and entire +
Ariz. +  and N.Mex. +
2200-2400 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
spatulate +, linear +  and narrowly triangular +
0.52 cm5.2 mm <br />0.0052 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
2-valved +  and capsular +
compressed +  and ovate +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
latiseptate +, stipitate +, , +, subsessile +  and sessile +
angustiseptate +, terete +, not +, inflated +, not torulose +, obovoid +, ovoid +, ovate +, obpyriform +, oblong +, obdeltate +, obcordate +, lanceolate +, elliptic +, ellipsoid +, suborbicular +, orbicular +, subglobose +  and globose +
latiseptate +  and unsegmented +
angustiseptate +  and terete +
Pinyon-juniper communities on nearly barren outcrops of Todilto Limestone +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
sinuate;dentate;sinuate;dentate +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
purple +, white +, orange +  and yellow +
rudimentary +
obdeltate +, oblanceolate +, ovate +, obovate +  and spatulate +
Flowering May-early Jun. +
trinucleate +  and 3(-11)-colpate +
ascending +, spreading +, horizontal +  and erect +
slender +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
bifurcate +
rounded +  and narrowly oblong +
not +  and mucilaginous +
margined +  and winged +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
flattened +
persistent +  and caducous +
distinct +
spreading +  and erect +
not saccate +, subsaccate +, deltate +, ovate +, oblong +, elliptic +, lanceolate +  and linear +
perforate +  and complete +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
prostrate +  and decumbent +
2-lobed +  and entire +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Coulterina +  and Lesquerella +
Physaria navajoensis +
Physaria +
species +
simple +, stalked +, subsessile +  and sessile +
stellate-scalelike +
coiled +, 1-7-veined +  and veined +
anastomosing +
perennial +, biennial +  and annual +
aquatics +  and terrestrial +
glabrous +  and pubescent +
15 +, 12 +, 10 +, 9 +, 8 +, 7 +, 6 +, 5 +  and 4 +