Piperia elegans subsp. elegans

IllustratedEndemic
Basionym: Habenaria maritima (Sprengel) S. Watson 1892
Synonyms: Habenaria elegans var. maritima (Greene) Ames Habenaria elegans var. multiflora (Rydberg) Peck Habenaria greenei Greene Habenaria unalascensis subsp. maritima Habenaria unalascensis var. maritima (Greene) Correll Piperia maritima Piperia multiflora
Treatment appears in FNA Volume 26. Treatment on page 573. Mentioned on page 570, 572, 574.

Plants 12–73 (–100) cm. Flowers pleasantly scented or musky; petals and lip pale green to white; spur 7–14 mm; viscidia oblong, 0.6–1 mm, usually more than 2 times as long as wide.


Phenology: Flowering Jun–Sep/Oct.
Habitat: Conifer and mixed evergreen forests, coastal scrub, exposed coastal bluff
Elevation: 0–1700 m

Distribution

V26 1170-distribution-map.jpg

B.C., Calif., Idaho, Mont., Oreg., Wash.

Discussion

Plants on exposed coastal bluffs are shorter and stouter than plants in sheltered or inland localities. Plants in warmer interior locations flower earlier than those on the coast and have fewer stem bracts. The flowers are pollinated by noctuid moths (J. D. Ackerman 1977).

A population in San Francisco, California, is also unusual by having basally attenuate stems and by flowering very late (Sep–Oct).

Selected References

None.

Lower Taxa

None.
self-pollination +
James D. Ackerman +  and Randall Morgan +
(Lindley) Rydberg +
Habenaria maritima +
indehiscent +
simple +  and sessile +
conduplicate +  and plicate +
20 cm200 mm <br />0.2 m <br /> (38 cm380 mm <br />0.38 m <br />) +
6 cm60 mm <br />0.06 m <br /> (20 cm200 mm <br />0.2 m <br />) +
lanceolate +  and oblanceolate +
1 cm10 mm <br />0.01 m <br /> (9 cm90 mm <br />0.09 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (2.4 cm24 mm <br />0.024 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (?) +  and 1.2 cm12 mm <br />0.012 m <br /> (?) +
inconspicuous +
parallel +  and close +
B.C. +, Calif. +, Idaho +, Mont. +, Oreg. +  and Wash. +
0–1700 m +
musky +  and scented +
ellipsoid;sausage-shaped +
Conifer and mixed evergreen forests, coastal scrub, exposed coastal bluff +
lateral +  and terminal +
2.5 cm25 mm <br />0.025 m <br /> (40 cm400 mm <br />0.4 m <br />) +
ovate-elliptic +  and oblong-lanceolate +
petiolate +, sessile +  and simple +
duplicate +, convolute +, whorled +, opposite +, distichous +  and alternate +
ephemeral +
cauline +  and basal +
2 (?) +  and 6 (?) +
reduced +
pale green +  and white +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
triangular-lanceolate +  and lanceolate +
larger +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
differing in form and color +
1-many-flowered +  and cymose +
pendent +  and erect +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
pale green +  and white +
straight +
recurved +  and spreading +
ovate;oblong-lanceolate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
Flowering Jun–Sep/Oct. +
Bull. Torrey Bot. Club +
spicate +  and many-flowered +
aerial +  and subterranean +
cinnamon brown +
minute +
free +, distinct +, adnate +  and connate +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
foliaceous +
not +  and alike +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Illustrated +  and Endemic +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
clavate +  and filiform +
0.7 cm7 mm <br />0.007 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
pendent +  and erect +
proliferous +
swollen +
thickened +  and stout +
Habenaria elegans var. maritima +, Habenaria elegans var. multiflora +, Habenaria greenei +, Habenaria unalascensis subsp. maritima +, Habenaria unalascensis var. maritima +, Piperia maritima +  and Piperia multiflora +
Piperia elegans subsp. elegans +
Piperia elegans +
subspecies +
ellipsoid-ovoid +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.015 cm0.15 mm <br />1.5e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
73 cm730 mm <br />0.73 m <br /> (100 cm1,000 mm <br />1 m <br />) +
aquatic +, lithophytic +, terrestrial +  and epiphytic +
saprophytic +, photosynthetic +  and mycotrophic +