Piperia unalascensis

(Sprengel) Rydberg

Bull. Torrey Bot. Club 28: 270. 1901.

IllustratedEndemic
Basionym: Spiranthes unalascensis Sprengel Syst. Veg. 3: 708. 1826
Synonyms: Habenaria schischmareffiana Chamisso Habenaria unalaschensis (Sprengel) S. Watson Platanthera foetida Geyer ex Hooker Platanthera unalaschcensis (Sprengel) Kurtz
Treatment appears in FNA Volume 26. Treatment on page 575. Mentioned on page 573, 574, 576.

Plants 9–70 cm. Stems swollen to uniform diameter toward base, 0.7–6.1 mm diam. distal to leaves; bracts 1–8. Leaves prostrate; blade 5–16 (–20) × 1.3–3.5 cm. Inflorescences usually sparsely flowered, 3–44 cm; rachis usually longer than peduncle; bracts 3–23 mm. Flowers ± translucent green, fragrance nocturnal but lingering during day, musky or soapy, sometimes honeylike; sepals ± translucent, 2–4.2 × 1–2.6 mm; dorsal sepal ovate to oblong; lateral sepals spreading to strongly recurved; petals usually projecting to erect, ± straight-sided, asymmetrically ovate to linearlanceolate, 2–5.5 × 0.6–2 mm; lip ± deflexed, broadly ovate to lanceolate-elliptic, 2–5 × 1–3 mm, apex usually somewhat upcurved; spur horizontal to decurved, 2–5.5 mm, ± equal to lip; viscidia broadly elliptic-ovate, 0.15–0.5 × 0.1–0.3 mm; rostellum blunt. Capsules 3.5–10.5 mm. Seeds tan to cinnamon brown.


Phenology: Flowering Jun–Aug; Apr–May, lowlands of n Calif.
Habitat: Coniferous and mixed evergreen forests, generally dry sites, rarely coastal bluffs
Elevation: 0–3000 m

Distribution

V26 1175-distribution-map.jpg

Alta., B.C., Nfld. and Labr. (Nfld.), Ont., Que., Alaska, Calif., Colo., Idaho, Mich., Mont., Nev., N.Mex., Oreg., S.Dak., Utah, Wash., Wyo.

Discussion

The racemes in Piperia unalascensis are usually slender and sparsely flowered; racemes of uncommon coastal populations (including the type) are short, stout, and densely flowered. Plants of the coast ranges and the Pacific Northwest are stouter and have broader sepals and petals than do interior and montane forms. Two sympatric forms appear to be in the Sierra Nevada, differing in lip morphology and scent.

Selected References

None.

Lower Taxa

None.
... more about "Piperia unalascensis"
self-pollination +
James D. Ackerman +  and Randall Morgan +
(Sprengel) Rydberg +
Spiranthes unalascensis +
indehiscent +
simple +  and sessile +
conduplicate +  and plicate +
16 cm160 mm <br />0.16 m <br /> (20 cm200 mm <br />0.2 m <br />) +
5 cm50 mm <br />0.05 m <br /> (16 cm160 mm <br />0.16 m <br />) +
lanceolate +  and oblanceolate +
1.3 cm13 mm <br />0.013 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
0.35 cm3.5 mm <br />0.0035 m <br /> (?) +  and 1.05 cm10.5 mm <br />0.0105 m <br /> (?) +
inconspicuous +
parallel +  and close +
Alta. +, B.C. +, Nfld. and Labr. (Nfld.) +, Ont. +, Que. +, Alaska +, Calif. +, Colo. +, Idaho +, Mich. +, Mont. +, Nev. +, N.Mex. +, Oreg. +, S.Dak. +, Utah +, Wash. +  and Wyo. +
0–3000 m +
translucent green +
honeylike +
ellipsoid;sausage-shaped +
Coniferous and mixed evergreen forests, generally dry sites, rarely coastal bluffs +
lateral +  and terminal +
3 cm30 mm <br />0.03 m <br /> (44 cm440 mm <br />0.44 m <br />) +
spreading +  and strongly recurved +
petiolate +, sessile +  and simple +
duplicate +, convolute +, whorled +, opposite +, distichous +  and alternate +
ephemeral +
cauline +  and basal +
2 (?) +  and 6 (?) +
reduced +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
broadly ovate +  and lanceolate-elliptic +
larger +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
differing in form and color +
1-many-flowered +  and cymose +
pendent +  and erect +
straight-sided +
2mm;5.5mm +
usually projecting +  and erect +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Flowering Jun–Aug +  and Apr–May, lowlands of n Calif. +
Bull. Torrey Bot. Club +
spicate +  and many-flowered +
usually longer +
aerial +  and subterranean +
tan +  and cinnamon brown +
minute +
free +, distinct +, adnate +  and connate +
foliaceous +
not +  and alike +
0.1 cm1 mm <br />0.001 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
Illustrated +  and Endemic +
horizontal +  and decurved +
clavate +  and filiform +
0.2 cm2 mm <br />0.002 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.61 cm6.1 mm <br />0.0061 m <br />) +
pendent +  and erect +
proliferous +
swollen +
thickened +  and stout +
Habenaria schischmareffiana +, Habenaria unalaschensis +, Platanthera foetida +  and Platanthera unalaschcensis +
Piperia unalascensis +
species +
ellipsoid-ovoid +
0.015 cm0.15 mm <br />1.5e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
elliptic-ovate +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
aquatic +, lithophytic +, terrestrial +  and epiphytic +
saprophytic +, photosynthetic +  and mycotrophic +