Polystichum scopulinum

(D. C. Eaton) Maxon

Fern Bull. 8: 29. 1900.

Common names: Rock sword fern polystic des rochers
Endemic
Basionym: Aspidium aculeatum var. scopulinum D. C. Eaton Ferns N. Amer. 2: 125. 1880
Synonyms: Polystichum mohrioides var. scopulinum (D. C. Eaton) Fernald
Treatment appears in FNA Volume 2.

Stems ascending. Leaves erect, 1–3 (–5) dm; bulblets absent. Petiole 1/5–1/3 length of leaf, densely scaly but scales falling off distally; scales light-brown, abruptly diminishing in size distally. Blade narrowly lanceolate, 1-pinnate-pinnatifid, base narrowed. Pinnae oblong-lanceolate, overlapping, folded inward and twisted horizontally, 1–3 cm; base oblique; margins serrulate with teeth curved inward; apex obtuse to cuspidate with subapical teeth smaller than apical tooth; microscales narrowly lanceolate, with stout projections, sparse, on abaxial surface only. Indusia entire-ciliate. Spores brown. 2n = 164.


Habitat: Rock crevices and at base of boulders, serpentine to acidic substrates, usually exposed to full sun
Elevation: 0–3500 m

Distribution

V2 751-distribution-map.gif

B.C., Nfld. and Labr. (Nfld.), Que., Ariz., Calif., Colo., Idaho, Mont., Nev., Oreg., Utah, Wash., Wyo.

Discussion

Polystichum scopulinum is widely distributed in the United States west of the 110th meridian, where it occurs in sporadic, usually small populations. The species is abundant only on montane serpentine outcrops. The populations in Newfoundland and Quebec are dramatically disjunct.

Polystichum scopulinum is an allopolyploid, believed on morphologic grounds to be derived from P. imbricans × lemmonii (D. H. Wagner 1979). Based on putative hybridization between P. scopulinum and P. munitum (P. S. Soltis et al. 1989; W. H. Wagner Jr. 1973), however, P. munitum may also be involved. This hybrid is discussed under P. californicum.

Selected References

None.

Lower Taxa

None.

"dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property."/5lengthofleaf" is not declared as a valid unit of measurement for this property.

... more about "Polystichum scopulinum"
obtuse +  and cuspidate +
smaller +
David H. Wagner +
(D. C. Eaton) Maxon +
narrowed +
Aspidium aculeatum var. scopulinum +
simple to commonly +
1-pinnate-pinnatifid;lanceolate +
reduced +
leathery;leathery +
roundish +
Rock sword fern +  and polystic des rochers +
B.C. +, Nfld. and Labr. (Nfld.) +, Que. +, Ariz. +, Calif. +, Colo. +, Idaho +, Mont. +, Nev. +, Oreg. +, Utah +, Wash. +  and Wyo. +
0–3500 m +
inflated +
often +  and glabrous +
Rock crevices and at base of boulders, serpentine to acidic substrates, usually exposed to full sun +
caducous +  and persistent +
monomorphic +
evergreen +
spinulose-toothed +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
sessile +  and short-petiolulate +
round +, cuplike +, hoodlike +, reniform +, falcate +  and linear +
2 +  and 3 +
light-brown +
persistent +
scaly +  and glabrous +
linear;lanceolate +
diminishing +
discrete +
reniform +  and oblong +
dictyostelic +, unbranched +  and branched +
Polystichum mohrioides var. scopulinum +
Polystichum scopulinum +
Polystichum +
species +
anastomosing +
epiphytic +  and hemiepiphytic +