Polytrichastrum papillatum

G. L. Smith

J. Hattori Bot. Lab. 38: 633, figs. 12–24. 1974,.

Basionym: Polytrichum papillatum (G. L. Smith) L. E. Anderson H. A. Crum & W. R. Buck
Treatment appears in FNA Volume 27. Treatment on page 128. Mentioned on page 125.

Plants slender, with the aspect of P. alpinum var. septentrionale. Stems to 2 cm, leafless below, leafy above. Leaves channeled and incurved when dry, spreading when moist, narrowly triangular, from a broadly elliptic, hyaline-margined sheathing base; marginal lamina serrate to almost entire; lamellae 5–6 cells high, crenate in profile, with each cell crowned by a papillose knob, the marginal cells in section pyriform. [Seta to 1.5 cm, curved. Capsule subspherical; exothecial cells with a distinct thin spot; peristome teeth ca. 40, hyaline, now and then compound.]


Habitat: Substrate and elevation undetermined

Distribution

V27 152-distribution-map.gif

Alaska, Asia (Kashmir), Asia (e Nepal)

Discussion

Polytrichastrum papillatum is known in North America from a single, sterile collection from southern Alaska (Valdez), collected by F. J. Hermann, but is possibly more widespread and overlooked. The marginal cells of the lamellae are coarsely papillose, and resemble those of P. alpinum in section. In profile, however, the lamellae are crenate, the marginal cells crowned by a papillose knob. The exothecial cells with a pale but distinct thin spot in the outer wall further distinguish this species from P. alpinum.

Selected References

None.

Lower Taxa

None.

"broadened" is not a number.

cucullate +
prominent +
Gary L. Smith Merrill +
G. L. Smith +
toothed +
hyaline-margined +
differentiated +
Polytrichum papillatum +
distinct +
abrupt +, crisped +  and undulate +
firm-textured +
not pitted +  and papillose +
monoicous +, dioicous +, sexual +  and vegetative +
crenate +
sparsely ciliate;smooth +
cucullate +
4-6-angled +  and terete +
short-excurrent +
prolonged +
toothed +
undifferentiated +
Alaska +, Asia (Kashmir) +  and Asia (e Nepal) +
scabrous +, mammillose +  and smooth +
Substrate and elevation undetermined +
cylindric +  and tapering +
papillose +
photosynthetic +
5 +  and 6 +
serrate +, crenulate +  and serrulate +
spreading +  and incurved +
triangular +  and channeled +
membranous +
1(-3)-stratose +
papillose +  and smooth +
crenulate +  and serrulate +
shape +  and size +
polygamous +, dioicous +  and sexual +
marginal lamina +  and almost entire +
sharply toothed +  and entire +
interwoven +
toothlike +
J. Hattori Bot. Lab. +
underground +
undifferentiated +
hyaline-margined +
differentiated +
larger +  and minute +
leafy +  and leafless +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Polytrichastrum papillatum +
Polytrichastrum +
species +
unjointed +
not keeled +