Pseudognaphalium beneolens

(Davidson) Anderberg

Opera Bot. 104: 147. 1991.

Common names: Fragrant rabbit-tobacco
Basionym: Gnaphalium beneolens Davidson Bull. S. Calif. Acad. Sci. 17: 17, unnumb. fig. p. 16. 1918
Synonyms: Gnaphalium canescens subsp. beneolens (Davidson) Stebbins & D. J. Keil Pseudognaphalium canescens subsp. beneolens (Davidson) Kartesz
Treatment appears in FNA Volume 19. Treatment on page 419. Mentioned on page 417.

Annuals or short-lived perennials, 30–80 (–110) cm; taprooted. Stems persistently tomentose, not glandular. Leaf-blades mostly linear, 3–6 cm × 1.5–3.5 mm (sometimes smaller distally), bases not clasping, decurrent 5–15 mm, margins flat, faces concolor, loosely tomentose, not glandular. Heads usually in loose, paniculiform arrays. Involucres turbinate-campanulate, 5–6 mm. Phyllaries in (4–) 5–6 (–7) series, white (opaque, dull to shiny), ovate to ovate-oblong (inner usually with filiform keel and slight apiculum), glabrous. Pistillate florets (39–) 44–69. Bisexual florets 5–8 (–11). Cypselae ridged, smooth or weakly papillate-roughened. 2n = 14.


Phenology: Flowering (Apr–)Jun–Oct.
Habitat: Dry, open slopes and ridges, streambeds, road banks and other disturbed sites, sandy flats, dunes, coastal sage scrub, chaparral, yellow pine, foothill pine, blue oak woodland
Elevation: (1–)50–800(–2000) m

Distribution

V19-684-distribution-map.gif

Calif., Mexico (Baja California)

Discussion

Pseudognaphalium beneolens differs from P. thermale in its leaves linear throughout, heads usually in elongate, paniculiform arrays, larger heads (greater numbers of phyllaries in greater numbers of series) with phyllaries more opaque and duller, and greater numbers of bisexual florets. The cauline leaves of P. beneolens tend to become curving-coiling. In areas of sympatry, habitats of P. beneolens are characteristically at lower elevations than those of P. thermale.

Selected References

None.

Lower Taxa

None.
white +  and gray +
tomentose +  and velutinous +
grayish;greenish +
arachnose +, glabrescent +  and glabrous +
truncate-penicillate;truncate +
scarious +
mostly ovate +  and lanceovate or linear +
paniculiform +
indeterminate +  and determinate +
Guy L. Nesom +
(Davidson) Anderberg +
decurrent +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
Gnaphalium beneolens +
compound +  and simple +
mostly narrowly lanceolate;oblanceolate +
5-ribbed +, 3-ribbed +  and 2-ribbed +
muriculate +, papillate +  and smooth +
Fragrant rabbit-tobacco +
actinomorphic +
papillate-roughened +  and smooth +
Calif. +  and Mexico (Baja California) +
(1–)50–800(–2000) m +
papillate-roughened +
winged +  and beaked +
dispersed +
Dry, open slopes and ridges, streambeds, road banks and other disturbed sites, sandy flats, dunes, coastal sage scrub, chaparral, yellow pine, foothill pine, blue oak woodland +
singly +  and aggregated +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
turbinate-campanulate +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
3 cm30 mm <br />0.03 m <br /> (6 cm60 mm <br />0.06 m <br />) +
1.5mm;3.5mm +
cauline +  and basal +
rarely lanceovate +  and lanceolate +
2-carpellate +
persistent +
Flowering (Apr–)Jun–Oct. +
bearing subulate enations +, hairy +  and bristly +
conic;more or less columnar +
4 +  and 6 +
exalbuminous +
decumbent +  and procumbent +
appendaged +  and truncate +
papillate +
Gnaphalium canescens subsp. beneolens +  and Pseudognaphalium canescens subsp. beneolens +
Pseudognaphalium beneolens +
Pseudognaphalium +
species +
80 cm800 mm <br />0.8 m <br /> (110 cm1,100 mm <br />1.1 m <br />) +
perennial +, short-lived +  and annual +
30 cm300 mm <br />0.3 m <br /> (80 cm800 mm <br />0.8 m <br />) +
shrub +  and subshrub +