Ptychostomum neodamense

(Itzigsohn) J. R. Spence

Phytologia 87: 21. 2005.

Basionym: Bryum neodamense Itzigsohn in J. K. A. Müller, Syn. Musc. Frond. 1: 258. 1848
Treatment appears in FNA Volume 28. Treatment on page 162. Mentioned on page 156, 165, 171.

Plants in dense or open turfs, green, red-green, or yellow-green. Stems 2–4 (–6) cm, tufted, comose, innovations elongate and evenly foliate; often strongly radiculose. Leaves yellow-green to green, proximal leaves distinctly black-gray with age, crowded, strongly twisted to contorted when dry, ovate, strongly concave, (1–) 2–3 (–4) mm, not much enlarged toward stem apex; base wide, not or weakly decurrent; margins revolute to mid leaf or beyond, limbidium strong, in 2 or 3 rows; apex broadly acute to obtuse; costa not reaching apex to percurrent, rarely short-excurrent, awn smooth; proximal laminal cells 3–4: 1, same width or wider than more distal cells; medial and distal cells rhomboidal, 14–20 µm wide, 2–3: 1, walls usually firm to incrassate. Specialized asexual reproduction absent. Sexual condition dioicous. Seta 1–3 (–4) cm. Capsule brown, elongate-ovate, symmetric, 3–5 mm, mouth yellow; operculum conic, apiculate; peristome well developed; exostome teeth yellow basally, hyaline distally, lamellae usually straight mid-tooth, pores absent along mid line; endostome not adherent to exostome, basal membrane high, 1/2 exostome height, segments with ovate perforations, cilia long, appendiculate. Spores (10–) 12–16 µm, finely papillose, pale-yellow or green.


Phenology: Capsules mature Jul–Aug.
Habitat: Wet soil, soil over rock, often calcareous
Elevation: low to high elevations (0-3000 m)

Distribution

V28 257-distribution-map.gif

Greenland, Alta., B.C., Nfld. and Labr., N.W.T., Nunavut, Ont., Yukon, Alaska, Calif., Colo., Eurasia

Discussion

Ptychostomum neodamense is closely related to P. bimum, P. subneodamense, and P. pseudotriquetrum, differing in its ovate, blunt, mostly non-decurrent leaves crowded along the stem. The costal awns are stout. D. T. Holyoak and L. Hedenäs (2006) did not consider P. neodamense a good species, based on a very limited sampling from Europe, but their results could also be interpreted to suggest that it may be distinct. Disjunct material from California closely matches European collections, but Colorado material differs somewhat in overall habit and ecology.

Selected References

None.

Lower Taxa

None.

"narrower" is not a number."narrow" is not a number.

... more about "Ptychostomum neodamense"
John R. Spence +
(Itzigsohn) J. R. Spence +
short-excurrent +  and percurrent +
denticulate +
reduced +
curved +  and straight +
decurrent +
Bryum neodamense +
cucullate +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
nutant +, inclined +  and suberect +
long-exserted +
elongate-ovate +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
inflated +
not differentiated +
appendiculate +
usually percurrent +  and long-excurrent +
subpercurrent +
rhomboidal;hexagonal +
Greenland +, Alta. +, B.C. +, Nfld. and Labr. +, N.W.T. +, Nunavut +, Ont. +, Yukon +, Alaska +, Calif. +, Colo. +  and Eurasia +
low to high elevations (0-3000 m) +
hyaline;pale-yellow +
not adherent +
hyaline +, brown +, orange +  and yellow +
hyaline +  and yellow +
Wet soil, soil over rock, often calcareous +
not +  and differentiated +
narrowly ovatelanceolate;triangular +
polyoicous +, monoicous +, dioicous +  and sexual +
imbricate;variously contorted +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
yellow-green +  and green +
ovatelanceolate +  and ovate +
shrunken +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
2-stratose +  and 1-stratose +
revolute entire +  and denticulate +
1 (?) +, 2 (?) +  and 3 (?) +
short-rectangular +  and quadrate +
lateral +  and terminal +
lateral +  and terminal +
diplolepidous-alternate +
Capsules mature Jul–Aug. +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
differentiated +
smooth;papillose +
not absent +
3 cm30 mm <br />0.03 m <br /> (4 cm40 mm <br />0.04 m <br />) +
straight +  and flexuose +
elongate +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
green +  and pale-yellow +
germinating +
variable +
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Ptychostomum neodamense +
Ptychostomum subg. Cladodium +
species +
lanceolate +
distinct +
not +  and porose +
straight +
specialized +
dioicous +, sexual +  and asexual +
sinuate +
incrassate +
yellow-green +, red-green +  and green +
large;small to very +