Ptychostomum pendulum

Hornschuch

Syll. Pl. Nov. 1: 64. 1822.

Synonyms: Bryum algovicum Sendtner ex Müller Hal.
Treatment appears in FNA Volume 28. Treatment on page 163. Mentioned on page 157, 164.

Plants in dense or open turfs, green, red-green, or yellow-green. Stems 0.5–1 (–1.5) cm, tufted, comose, innovations elongate and evenly foliate, often strongly radiculose. Leaves yellow-green to red-green, weakly to strongly contorted or shrunken when dry, ovatelanceolate, concave, (0.4–) 0.8–1.5 (–2.5) mm, enlarged toward stem apex; base not decurrent; margins revolute to mid leaf, limbidium strong, in 2 or 3 rows; apex acuminate; costa long-excurrent, awn denticulate; proximal laminal cells 3–4: 1, same width or narrower than more distal cells; medial and distal cells rhomboidal, 12–18 µm wide, 3–4: 1, walls thin. Specialized asexual reproduction absent. Sexual condition synoicous. Seta 1–3 cm. Capsule brown, pyriform, symmetric, 2–3 mm, mouth red; operculum short-convex, apiculate; peristome reduced; exostome teeth yellow basally, hyaline distally, appearing chambered due to adherence of endostome, lamellae indistinct, pores absent along mid line; endostome strongly adherent to exostome, basal membrane 1/3–1/2 exostome height, segments with ovate perforations, cilia reduced to nearly absent. Spores with size variable in same collections and capsules, (22–) 25–30 (–36) µm, strongly papillose, pale-brown.


Phenology: Capsules mature May–Aug.
Habitat: Dry calcareous soil or rock
Elevation: low to high elevations (0-4000 m)

Distribution

V28 260-distribution-map.gif

Greenland, Alta., B.C., Man., Nfld. and Labr., N.W.T., N.S., Nunavut, Ont., Que., Sask., Yukon, Alaska, Ariz., Calif., Colo., Conn., D.C., Idaho, Ill., Ind., Kans., Maine, Mass., Mich., Minn., Mont., Nebr., Nev., N.H., N.J., N.Dak., Ohio, Oreg., Pa., S.Dak., Tenn., Tex., Utah, Vt., Wash., W.Va., Wis., Wyo., South America, Eurasia, n Africa, Pacific Islands (New Zealand), Subantarctic Islands

Discussion

Ptychostomum pendulum is a small plant found in dry sites, similar overall to P. archangelicum and P. inclinatum. The larger spores and unusual chambered appearance of the exostome teeth are diagnostic. Capsules are needed for identification.

Selected References

None.

Lower Taxa

None.

"narrower" is not a number."narrow" is not a number.

... more about "Ptychostomum pendulum"
acuminate +
John R. Spence +
Hornschuch +
denticulate +
reduced +
curved +  and straight +
not decurrent +
Undefined subg. Cladodium +
cucullate +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
nutant +, inclined +  and suberect +
long-exserted +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
inflated +
not differentiated +
appendiculate +
reduced +
long-excurrent +
subpercurrent +
rhomboidal;hexagonal +
Greenland +, Alta. +, B.C. +, Man. +, Nfld. and Labr. +, N.W.T. +, N.S. +, Nunavut +, Ont. +, Que. +, Sask. +, Yukon +, Alaska +, Ariz. +, Calif. +, Colo. +, Conn. +, D.C. +, Idaho +, Ill. +, Ind. +, Kans. +, Maine +, Mass. +, Mich. +, Minn. +, Mont. +, Nebr. +, Nev. +, N.H. +, N.J. +, N.Dak. +, Ohio +, Oreg. +, Pa. +, S.Dak. +, Tenn. +, Tex. +, Utah +, Vt. +, Wash. +, W.Va. +, Wis. +, Wyo. +, South America +, Eurasia +, n Africa +, Pacific Islands (New Zealand) +  and Subantarctic Islands +
low to high elevations (0-4000 m) +
hyaline;pale-yellow +
hyaline +  and yellow +
hyaline +, brown +, orange +  and yellow +
Dry calcareous soil or rock +
not +  and differentiated +
narrowly ovatelanceolate;triangular +
polyoicous +, monoicous +, dioicous +  and sexual +
imbricate;variously contorted +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
yellow-green +  and red-green +
erect;erect-spreading +
concave +  and ovatelanceolate +
enlarged +  and shrunken +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
2-stratose +  and 1-stratose +
revolute entire +  and denticulate +
1 (?) +, 3 (?) +  and 4 (?) +
short-rectangular +  and quadrate +
short-convex +
lateral +  and terminal +
lateral +  and terminal +
diplolepidous-alternate +
Capsules mature May–Aug. +
Syll. Pl. Nov. +
differentiated +
smooth;papillose +
not absent +
straight +  and flexuose +
elongate +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
germinating +
variable +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Bryum algovicum +
Ptychostomum pendulum +
Ptychostomum subg. Cladodium +
species +
lanceolate +
distinct +
not +  and porose +
straight +
specialized +
synoicous +, sexual +  and asexual +
sinuate +
incrassate +
yellow-green +, red-green +  and green +
large;small to very +