Rhamnus utilis var. utilis

Introduced
Treatment appears in FNA Volume 12. Treatment on page 51.

Shrubs or small trees, 1–4 m, armed with thorns. Branchlets brown to purple-red, glabrate. Leaves deciduous, opposite to subopposite, sometimes fascicled on short-shoots; petiole 5–15 mm; blade light green abaxially, glossy darker green adaxially, commonly yellowish or pale in age, oblong to elliptic, obovate-elliptic, or oblong-lanceolate, (4–) 6–11 (–13) cm, usually (2.5–) 2.8–3.5 (–3.8) times longer than wide, subcoriaceous, base cuneate, margins serrulate to crenate-serrate, apex acute, abaxial surface puberulent along veins or at least in vein-axils, hairs wavy or curled, yellow, adaxial surface glabrous; secondary-veins (4–) 5–7 (–8) pairs, all diverging at nearly same angle. Inflorescences fascicles. Pedicels 5–10 mm. Sepals 4. Petals 4. Drupes black, globose to subglobose, 5–7 mm; stones 2.


Phenology: Flowering Apr–Jun.
Habitat: Old fields, thickets, meadows, degraded woods, woods edges, commonly with Rhamnus cathartica and R. davurica.
Elevation: 200–300 m.

Distribution

V12 242-distribution-map.jpg

Introduced; Conn., Ill., Iowa, Mich., Ohio, Asia (China), Asia (Japan), Asia (Korea)

Discussion

According to E. G. Voss (1972–1996, vol. 2), some plants of Rhamnus utilis in Michigan are similar to forms of R. davurica and perhaps represent hybrids between the two. Hybridization between R. cathartica and R. utilis has been documented (N. L. Gil-Ad and A. A. Reznicek 1997).

Selected References

None.

Lower Taxa

None.

"thin" is not a number.

perigynous +  and epigynous +
Guy L. Nesom +  and John O. Sawyer† +
Decaisne +
cuneate +
11 cm110 mm <br />0.11 m <br /> (13 cm130 mm <br />0.13 m <br />) +
pale +, yellowish +, darker green +  and light green +
(2.5-)2.8-3.5(-3.8) +
3[-5]-veined +  and pinnate +
spinulose +, spinose +, serrate +  and entire +
denticulate +, dentate +, crenulate +, crenate +  and serrulate +
oblong;elliptic obovate-elliptic or oblong-lanceolate +
6 cm60 mm <br />0.06 m <br /> (11 cm110 mm <br />0.11 m <br />) +
subcoriaceous +
Conn. +, Ill. +, Iowa +, Mich. +, Ohio +, Asia (China) +, Asia (Japan) +  and Asia (Korea) +
globose +  and subglobose +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
200–300 m. +
bisexual +  and unisexual +
Old fields, thickets, meadows, degraded woods, woods edges, commonly with Rhamnus cathartica and R. davurica. +
curled +  and wavy +
circumscissile +
free +  and adnate +
campanulate +  and cupulate +
unisexual +  and bisexual +
opposite +  and subopposite +
deciduous +
serrulate to crenate-serrate +
intrastaminal +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
not fleshy +
perigynous +  and epigynous +
yellowish +  and cream +
adnate +  and distinct +
not clawed +  and spatulate +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
Flowering Apr–Jun. +
2-4-carpellate +
Compt. Rend. Hebd. Séances Acad. Sci. +
oblong-obovoid +  and obovoid +
greenish white +  and white +
distinct +
crested;not keeled;ovate-triangular +
opposite +  and subopposite +
Introduced +
4 +  and 5 +
2 +  and 4 +
Rhamnus utilis var. utilis +
Rhamnus utilis +
variety +
deciduous +  and evergreen +
tree +  and shrub +
spreading +  and erect +
polygamous +, dioecious +  and synoecious +