Sagina maxima subsp. crassicaulis

(S. Watson) G. E. Crow

Rhodora 80: 79. 1978.

Illustrated
Basionyms: Sagina crassicaulis S. Watson Proc. Amer. Acad. Arts 18: 191. 1883
Treatment appears in FNA Volume 5. Treatment on page 147. Mentioned on page 146.

Plants perennial, glabrous or mostly so. Stems spreading, decumbent, or procumbent, glabrous, nodes frequently purple-tinged. Leaves: basal leaves in rosette of broadly linear, fleshy leaves, or absent with primary or secondary tufts of ascending, linear basal leaves, these usually less fleshy than rosette leaves (rosettes rarely present in plants occurring north of Washington); cauline leaf-blades: proximal 6–15 mm, distal 3–5 mm, glabrous. Pedicels slender to stout, glabrous. Flowers: calyx glabrous; sepals ovate to nearly orbiculate, (2–) 2.5–3 (–3.5) mm; petals elliptic to orbiculate, (1.5–) 2–2.5 (–3) mm, slightly shorter than sepals. Capsules (3–) 3.5–4 (–4.5) mm. Seeds smooth to slightly pebbled. 2n = 46, 66.


Phenology: Flowering spring–early autumn.
Habitat: Coastal, moist, sandy bluffs, crevices of rock cliffs, at or near high-tide mark, gravelly-sandy beaches
Elevation: 0-10 m

Distribution

V5 302-distribution-map.gif

B.C., Alaska, Calif., Oreg., Wash., Asia (Kamchatka)

Discussion

Integradation occurs where the range of subsp. crassicaulis overlaps with that of subsp. maxima. Variation of pubescence in populations on Vancouver Island and the Queen Charlotte Islands ranges from completely glabrous specimens typical of subsp. crassicaulis to individuals with pedicels and calyx bases weakly pubescent, to others with densely pubescent pedicels. Subspecies crassicaulis is far more common than subsp. maxima.

Selected References

None.

Lower Taxa

None.

"/4" is not declared as a valid unit of measurement for this property.

glandular-pubescent +  and glabrous +
Garrett E. Crow +
(S. Watson) G. E. Crow +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
Sagina crassicaulis +
Laciniate (?) +, Emarginate (?) +, Jagged (?) +, 2-fid (?) +  and Entire (?) +
linear +  and subulate +
reduced +
herbaceous +  and scarious +
glandular-pubescent +  and glabrous +
0.4 cm4 mm <br />0.004 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
1/4 +  and 1/2 +
globose;ovoid +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
minute +
shallow +
conspicuous +
scarious +
dish--shaped +
expanded +
0.7 cm7 mm <br />0.007 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
procumbent +  and decumbent +
glandular-pubescent +
B.C. +, Alaska +, Calif. +, Oreg. +, Wash. +  and Asia (Kamchatka) +
0-10 m +
straight +  and curved +
central +  and peripheral +
inconspicuous +
unisexual +  and bisexual +
stalked +  and simple +
Coastal, moist, sandy bluffs, crevices of rock cliffs, at or near high-tide mark, gravelly-sandy beaches +
disc--shaped +  and dish +
not stipulate +  and petiolate +
connate +
purple-tinged +, whitish +  and hyaline +
purple-tinged +
swollen +
3 +  and 5 +
crassinucellate +  and bitegmic +
campylotropous +
spreading;erect +
slender +  and stout +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
fugacious +
5 +  and 4 +
0.2 cm2 mm <br />0.002 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
Flowering spring–early autumn. +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
smooth +  and slightly pebbled +
reniform +
0.3 cm3 mm <br />0.003 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
persistent +
distinct +
5 +  and 4 +
ovate;nearly orbiculate +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Illustrated +
branched +  and simple +
procumbent +  and decumbent +
terete;slightly angular +
subterminal +
[30 +, 5 +  and 4 +
papillate +
distinct +
5 (?) +  and 4 (?) +
list +  and count +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Spergella +
Sagina maxima subsp. crassicaulis +
Sagina maxima +
subspecies +
divided +
usually less fleshy +
10 +  and 8 +
rhizomatous +  and taprooted +
mostly +  and glabrous +
11 +, 7 +  and 6 +