Smelowskia borealis

(Greene) W. H. Drury & Rollins

Rhodora 54: 111. 1952.

Endemic
Basionym: Melanidion boreale Greene Ottawa Naturalist 25: 146. 1912
Synonyms: Acroschizocarpus kolianus Gombócz Ermania borealis (Greene) Hultén Smelowskia borealis var. jordalii W. H. Drury & Rollins Smelowskia borealis var. koliana (Gombócz) W. H. Drury & Rollins Smelowskia borealis var. villosa W. H. Drury & Rollins
Treatment appears in FNA Volume 7. Treatment on page 673. Mentioned on page 672, 674.

Plants sometimes canescent basally; caudex usually simple, rarely branched. Stems: several from base, often unbranched or branched proximally, 0.6–3 (–3.5) dm, trichomes simple, to 1 mm, mixed with smaller, dendritic ones. Basal leaves: petiole 0.5–3.2 cm, ciliate or not; blade obovate to ovate in outline, (terminal segments linear, oblong, or ovate), 0.7–2.5 cm × 4–14 mm, (terminal segments 0.2–1.1 cm × 1–5 mm), margins palmately (3 or) 5 (or 7) -lobed, (surfaces densely pubescent, trichomes dendritic, often mixed with larger, simple ones, to 1 mm), apex obtuse or subacute. Cauline leaves shortly petiolate or sessile; blade often similar to basal, smaller distally, margins sometimes pinnatifid. Racemes considerably elongated in fruit. Fruiting pedicels spreading to divaricate-ascending, (secund, often curved), proximalmost bracteate, 6–15 (–20) mm, pubescent, trichomes simple mixed with smaller, dendritic ones. Flowers: sepals (usually persistent), 2.5–3.2 mm; petals purple to lavender, suborbicular to obovate, 4–5 × 1.5–2 mm, narrowed to claw, 1.5–2.5 mm, apex rounded; anthers oblong, 0.4–0.5 mm. Fruits spreading to ascending, usually oblanceolate to spatulate or linear-oblanceolate, rarely oblong-obovate, angustiseptate, (8–) 12–28 × 4–8 mm, base cuneate, apex obtuse; valves each with prominent midvein and distinct lateral-veins, (rarely sparsely pubescent); ovules 10–18 per ovary; style 0.1–1.1 mm. Seeds 1.5–2.2 × 1–1.3 mm. 2n = 12.


Phenology: Flowering Jun–Aug.
Habitat: Loose talus, metamorphic slide rock, rocky slopes, scree, shale splinters, limestone rubble, alpine ridges, barren rocks, unstable talus in alpine glacial bowl
Elevation: 600-1700 m

Discussion

W. H. Drury Jr. and R. C. Rollins (1952) and Rollins (1993) divided Smelowskia borealis into four varieties based on the duration of sepals (persistent versus caducous), style length (less than versus greater than 0.5 mm), and density of indumentum. The various states of these three characters are present in all possible combinations, and they are not inherited in the combinations presented by these authors. Some forms of the species are densely villous and resemble S. johnsonii in their leaf indumentum and perhaps merit recognition as an infraspecific taxon (var. villosa). As delimited here, S. borealis is quite variable; more studies are needed to determine whether or not any of the four varieties merit recognition at some rank. Although numerous collections of the species were examined, none of the types was available for preparing this account. The species is easily distinguished from the remaining North American Smelowskia by having secund infructescences and distinctly angustiseptate fruits.

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Smelowskia borealis"
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
obtuse +, rounded +  and subacute +
Ihsan A. Al-Shehbaz +
(Greene) W. H. Drury & Rollins +
cuneate +
Melanidion boreale +
appendaged +  and unappendaged +
not auriculate +
0.7 cm7 mm <br />0.007 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
entire +, 2-pinnatisect +  and 1-pinnatisect +
dissected +, repand +, sinuate +, crenate +  and dentate +
obovate;ovate +
smaller +
0.4 cm4 mm <br />0.004 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
well-developed +
sessile +  and petiolate +
well-developed +
distinct +
differentiated +
emarginate +  and entire +
accumbent +  and incumbent +
N.W.T. +, Yukon +  and Alaska. +
600-1700 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.32 cm3.2 mm <br />0.0032 m <br />) +
stipitate +  and sessile +
0.8 cm8 mm <br />0.008 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
spreading +  and ascending +
angustiseptate +, oblong-obovate +, usually oblanceolate +  and spatulate or linear-oblanceolate +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Loose talus, metamorphic slide rock, rocky slopes, scree, shale splinters, limestone rubble, alpine ridges, barren rocks, unstable talus in alpine glacial bowl +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
pinnatifid;5-lobed +
smaller +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
purple +  and lavender +
rudimentary +
narrowed +, suborbicular +  and obovate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (3.2 cm32 mm <br />0.032 m <br />) +
Flowering Jun–Aug. +
trinucleate +  and 3(-11)-colpate +
suberect +, spreading +  and ascending +
slender +
mucilaginous +
aseriate +, biseriate +  and uniseriate +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
oblong-lanceolate +  and oblong +
0.1 cm1 mm <br />0.001 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
persistent +  and caducous +
distinct +
suberect +  and spreading +
perforated +  and complete +
reduced +
stipitate +  and sessile +
subterete +, terete +, angustiseptate +, 4-angled +, pyriform +, suboblong +, oblanceolate +, spatulate +, ellipsoid +, obovoid +, oblong +  and linear +
stipitate +  and sessile +
subterete +, terete +, angustiseptate +, 4-angled +, pyriform +, suboblong +, oblanceolate +, spatulate +, ellipsoid +, obovoid +, oblong +  and linear +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
erect +  and decumbent +
2-lobed +, entire +, conical +  and capitate +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
Acroschizocarpus kolianus +, Ermania borealis +, Smelowskia borealis var. jordalii +, Smelowskia borealis var. koliana +  and Smelowskia borealis var. villosa +
Smelowskia borealis +
Smelowskia +
species +
forked +  and stellate +
0 cm0 mm <br />0 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
coiled +, 1-7-veined +  and veined +
anastomosing +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
spreading +  and divaricate-ascending +
0.6 cm6 mm <br />0.006 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
aquatics +  and terrestrial +