Stanleya pinnata

(Pursh) Britton

Trans. New York Acad. Sci. 8: 62. 1889.

Endemic
Basionym: Cleome pinnata Pursh Fl. Amer. Sept. 2: 739. 1813
Treatment appears in FNA Volume 7. Treatment on page 697. Mentioned on page 695, 696.

Perennials, subshrubs, or shrubs; (sometimes suffrutescent); (glaucous or not), mostly glabrous. Stems erect, unbranched or branched (few) proximally and distally, (1.2–) 3–12 (–15.3) dm. Basal leaves (withered by flowering); similar to cauline. Cauline leaves: petiole 0.7–6.2 cm; blade (fleshy), oblanceolate to broadly lanceolate or ovate in outline, or (distally) lobed or linear to narrowly lanceolate, 3–15 cm (smaller distally), margins lyrate-pinnatifid or runcinate, or (distal) pinnately lobed or entire. Racemes somewhat dense. Fruiting pedicels horizontal to divaricate, 3–11 mm. Flowers: sepals oblong-linear, 8–16 mm; petals yellow, oblanceolate to oblong, 8–20 × 2–3 mm, claw 4–10 mm, wider at base, densely pubescent inside; filaments 11–28 mm, pilose at base; anthers 3–5 mm; gynophore 7–28 mm. Fruits usually spreading to divaricate, rarely ascending, often strongly curved downward, not tortuous, (smooth), terete, 3–9 cm × 1.5–3 mm; ovules 10–38 per ovary; style 0.2–0.6 mm. Seeds (sometimes black), oblong, 2.5–4.5 × 1.2–2 mm.

Distribution

V7 1202-distribution-map.gif

Ariz., Calif., Colo., Idaho, Kans., Mont., N.Dak., N.Mex., Nev., Oreg., S.Dak., Tex., Utah, Wyo.

Discussion

Varieties 3 (3 in the flora).

The three varieties of Stanleya pinnata are reasonably well-defined and can be easily separated from each other. Their maintenance here at varietal rank is tentative; a genus-wide phylogenetic study is much needed to establish whether or not they merit recognition as distinct species.

Key

1 Cauline leaf blades: margins (at least proximal ones) lyrate-pinnatifid to runcinate. Stanleya pinnata var. pinnata
1 Cauline leaf blades: margins usually entire, rarely proximalmost dentate > 2
2 Cauline leaf blades ovate to lanceolate; petals 11-16 mm, claws 6-9 mm; ovules 26-34 per ovary; gynophores pubescent basally; Colorado, Kansas, Nevada, Utah, Wyoming. Stanleya pinnata var. integrifolia
2 Cauline leaf blades linear-lanceolate; petals 8-9(-10) mm, claws 4-5 mm; ovules 10-20 per ovary; gynophores glabrous or nearly so; Texas. Stanleya pinnata var. texana

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Stanleya pinnata"
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Ihsan A. Al-Shehbaz +
(Pursh) Britton +
Cleome pinnata +
appendaged +  and unappendaged +
not +  and auriculate +
2-pinnatifid +, 1-pinnatifid +, lobed +  and entire +
much smaller +  and reduced +
urceolate +, campanulate +  and tubular +
branched +  and simple +
sessile +  and petiolate +
linear;narrowly lanceolate +
3 cm30 mm <br />0.03 m <br /> (15 cm150 mm <br />0.15 m <br />) +
well-developed +
distinct +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
differentiated +
emarginate +  and entire +
accumbent +  and incumbent +
Ariz. +, Calif. +, Colo. +, Idaho +, Kans. +, Mont. +, N.Dak. +, N.Mex. +, Nev. +, Oreg. +, S.Dak. +, Tex. +, Utah +  and Wyo. +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
1.1 cm11 mm <br />0.011 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
zygomorphic +  and actinomorphic +
latiseptate +  and long-stipitate +
not tortuous +  and curved +
3 cm30 mm <br />0.03 m <br /> (9 cm90 mm <br />0.09 m <br />) +
downward +, ascending +  and spreading +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
sessile +  and petiolate +
alternate +, not +  and rosulate +
basal +  and cauline +
connate +  and distinct +
decurrent +
dentate to pinnatifid +  and entire +
entire;lobed;runcinate;lyrate-pinnatifid;entire;lobed;runcinate;lyrate-pinnatifid +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
8mm;20mm +
rudimentary +
oblanceolate +  and oblong +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (6.2 cm62 mm <br />0.062 m <br />) +
trinucleate +  and 3(-11)-colpate +
Trans. New York Acad. Sci. +
slender +
0.3 cm3 mm <br />0.003 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
lichvar1983a +
mucilaginous +
aseriate +, uniseriate +  and biseriate +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
persistent +  and caducous +
distinct +
spreading +  and reflexed +
0.8 cm8 mm <br />0.008 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
entire +  and capitate +
2-lobed +  and entire +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
Cruciferae +
Stanleya pinnata +
Stanleya +
species +
dendritic +  and simple +
coiled +, 1-7-veined +  and veined +
anastomosing +
pubescent +  and glabrous +
perennial +, biennial +  and annual +
shrub +  and subshrub +
aquatics +  and terrestrial +