Syntrichia papillosissima

(Coppey) Loeske

Hedwigia 49: 44. 1910,.

Basionym: Barbula papillosissima Coppey Bull. Séances Soc. Sci. Nancy, sér. 3, 8: 314, plate 2, figs. f, g. 1908
Synonyms: Barbula ruralis var. hirsuta Venturi Tortula papillosissima Tortula ruralis var. hirsuta (Venturi) Paris
Treatment appears in FNA Volume 27. Treatment on page 626. Mentioned on page 619, 625, 627.

Stems 10–25 mm. Leaves clasping at base, infolded and twisted around the stem when dry, squarrose-recurved when moist, lingulate-ovate, 2.5–4 × 1–1.6 mm, canaliculate to keeled; margins tightly revolute in the proximal 3/4–7/8, entire; apices obtuse to acute; costa excurrent into a serrate, hyaline awn that often brown, sometimes broadly hyaline at base, strongly papillose abaxially and serrate because of projecting cell ends, yellowbrown; basal-cells abruptly differentiated, rectangular, 45–90 × 15–23 µm, quadrate to narrowly rectangular at the margins; distal cells quadrate to polygonal, 11–18 µm, with tall, bulging mammillae, bearing 1–2 papillae per cell, thick-walled and sometimes collenchymatous. Specialized asexual reproduction absent. Sexual condition dioicous. Seta brown, 12–18 mm. Capsule brown, 3–5 mm, curved, with an abrupt neck; operculum ca. 2 mm; peristome ca. 1.8 mm, the upper divisions twisted ca. 2 turns, yellowbrown, the basal membrane white, ca. 1/2 the total length. Spores 10–14 µm, papillose.


Habitat: Dry soil, rock
Elevation: moderate to high elevations

Distribution

V27 916-distribution-map.gif

B.C., Ariz., Calif., Colo., Idaho, Mont., Nev., N.Mex., Oreg., Utah, Wash., Mexico (Coahuila), Mexico (Nuevo León), s Europe, c Asia

Discussion

Syntrichia papillosissima is primarily a species of the Great Basin Desert north into the shrub-steppe ecosystems of the Columbia Basin, where it often occurs as a co-dominant with S. ruralis and S. caninervis. It is similar to a robust S. ruralis, differing most conspicuously in the extremely tall mammillae on the distal leaf cells, each crowned by only one or two papillae, unlike the shorter bulging cell surface bearing four or five papillae characterizing other species in the S. ruralis complex. Syntrichia papillosissima also has larger distal laminal cells, which are more pellucid than those of S. ruralis.

Selected References

None.

Lower Taxa

None.
... more about "Syntrichia papillosissima"
elongate +  and quadrate +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
apiculate +, emarginate +  and truncate +
Obtuse (?) +  and Acute (?) +
Brent D. Mishler +
(Coppey) Loeske +
brown +  and hyaline +
reniform +  and rounded +
quadrate +  and narrowly rectangular +
papillose +  and smooth +
usually slightly wider +
usually ovate;oblong +
Barbula papillosissima +
obovoid +
not differentiated +
brown +  and yellow +
acute +  and long-cucullate +
cleistocarpous +  and stegocarpous +
striolate +
cylindric +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
monoicous +, dioicous +  and sexual +
flat to bulging +
vesiculose +  and thick-walled +
Few (?) +  and Several (?) +
elongate-rhomboid +  and elongate +
tomentose +  and bare +
yellowbrown +  and hyaline +
papillose +
collenchymatous +  and thick-walled +
sometimes slightly longer than wide +
dioicous +, sexual +  and asexual +
quadrate +  and polygonal +
large +  and small +
B.C. +, Ariz. +, Calif. +, Colo. +, Idaho +, Mont. +, Nev. +, N.Mex. +, Oreg. +, Utah +, Wash. +, Mexico (Coahuila) +, Mexico (Nuevo León) +, s Europe +  and c Asia +
moderate to high elevations +
differentiated +
multicellular +
Dry soil, rock +
yellow +  and orange-red or red +
strong +  and weak +
1 +  and 3 +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
infolded +
reduced +
revolute +  and plane +
pale +  and colored +
distinct +
2 cm20 mm <br />0.02 m <br /> (?) +
photosynthetic +
compound +, simple +, hollow +  and solid +
1 +  and 2 +
flattened +  and 2-fid +
lateral +  and terminal +
blunt +, oblong-lingulate +  and ovate +
lateral +  and terminal +
1.8 cm18 mm <br />0.018 m <br /> (?) +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
synoicous +, autoicous +, dioicous +  and sexual +
cylindric +  and spherical +
straight +
elongate +
1.2 cm12 mm <br />0.012 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
2 +  and 1 +
brick-red +  and laminal +
2.5mm;4mm +
squarrose-recurved +
canaliculate +  and keeled +
1 cm10 mm <br />0.01 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
1mm +  and 1.6mm +
Barbula ruralis var. hirsuta +, Tortula papillosissima +  and Tortula ruralis var. hirsuta +
Syntrichia papillosissima +
Syntrichia +
species +
ovoid;cylindric +
yellow distally and red +, orange +, red +  and yellow +
stemless +  and perigoniate +
green;yellow or redbrown +
small +  and large +