Taraxia tanacetifolia

(Torrey & A. Gray) Piper

Contr. U.S. Natl. Herb. 11: 405. 1906.

Basionym: Oenothera tanacetifolia Torrey & A. Gray in War Department [U.S.] Pacif. Railr. Rep. 2(1): 121, plate 4. 1857
Synonyms: Camissonia tanacetifolia (Torrey & A. Gray) P. H. Raven C. tanacetifolia subsp. quadriperforata P. H. Raven Taraxia longiflora Nuttall ex Raimann T. tikurana A. Nelson
Treatment appears in FNA Volume 10.

Herbs usually densely, sometimes sparsely, short-hirtellous, hairs spreading or appressed, and/or sparselyto densely strigillose; taproot deep, woody, with numer­ous slender branches in age, producing new rosettes. Leaves 6.5–32 × 0.7–3.3 cm; petiole not winged, 1–8 cm; blade very narrowly elliptic, base attenuate to narrowly cuneate, margins deeply and irregularly pinnatifid, apex acute to long-acuminate. Flowers opening near sunrise; floral-tube 4–6.5 (–8.5) mm, soft-pilose in proximal 1/2 inside; sepals 5.5–13 mm; petals yellow, (8–) 10–23 mm; episepalous staminal filaments 5.5–12 mm, epipetalous ones 2.5–8 mm, anthers (2.3–) 2.8–3.5 mm; sterile prolongation of ovary 14–55 mm, style 9.5–20 (–25) mm, pilose near base, stigma exserted beyond anthers at anthesis. Capsules subterete, cylindric-ovoid, 7–25 × 3–5 mm, walls thick, evidently distended by seeds; sessile. Seeds uniformly tan to brown, curved-cylindric, 1.5–2 × 0.6–0.8 mm, pitted in rows. 2n = 14, 28, 42.


Phenology: Flowering May–Jul.
Habitat: Open areas on clay soil, moist swales and meadows, dry streambeds, edges of drying ponds.
Elevation: 400–2500 m.

Distribution

Calif., Idaho, Nev., Oreg., Wash.

Discussion

P. H. Raven (1969) determined that the majority of plants examined cytologically were tetraploid (2n = 28), making one diploid (2n = 14) determination. Raven also found hexaploid plants in a restricted area from Sierra Valley in Plumas and immediately adjacent Lassen and Sierra counties, California. They do not differ morphologically from diploid and tetraploid plants other than having greater than ten percent of pollen grains 4-pored, but there was no known diploid population that could have combined with tetraploids to give rise to this hexaploid. Raven named it Camissonia tanacetifolia subsp. quadriperforata; however, it is not currently recognized due to the lack of morphological differentiation. Raven also found Taraxia tanacetifolia to be primarily self-incompatible with possibly some self-compatible plants occurring.

Oenothera nuttallii Torrey & A. Gray 1840 (not Sweet 1830) pertains here.

Selected References

None.

Lower Taxa

None.

"thin" is not a number.

... more about "Taraxia tanacetifolia"
42 +, 28 +  and 14 +
0.23 cm2.3 mm <br />0.0023 m <br /> (0.28 cm2.8 mm <br />0.0028 m <br />) +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
acute +  and long-acuminate +
Warren L. Wagner +
(Torrey & A. Gray) Piper +
attenuate +  and narrowly cuneate +
Oenothera tanacetifolia +
indehiscent +  and loculicidal +
subentire +  and deeply sinuate or pinnatifid +
subsessile +  and thick-walled +
curved +  and straight +
0.7 cm7 mm <br />0.007 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
cylindric-ovoid +  and subterete +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Calif. +, Idaho +, Nev. +, Oreg. +  and Wash. +
400–2500 m. +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
basifixed +  and versatile +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
actinomorphic +
subsessile +  and thick-walled +
curved +  and straight +
indehiscent +  and dehiscent +
persistent +
tapering +, subterete +  and 4-angled cylindric-lanceoloid or ovoid or oblong-ellipsoid +
Open areas on clay soil, moist swales and meadows, dry streambeds, edges of drying ponds. +
appressed +  and spreading +
strigillose +
acaulescent +
perennial +
short-hirtellous +
whorled +, opposite +  and alternate +
deciduous +
6.5 cm65 mm <br />0.065 m <br /> (32 cm320 mm <br />0.32 m <br />) +
basal +  and cauline +
0.7 cm7 mm <br />0.007 m <br /> (3.3 cm33 mm <br />0.033 m <br />) +
abscission +
pinnatifid +
1.4 cm14 mm <br />0.014 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
numerous +  and 1 +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
1 cm10 mm <br />0.01 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
1 cm10 mm <br />0.01 m <br /> (8 cm80 mm <br />0.08 m <br />) +
Flowering May–Jul. +
0.4 cm4 mm <br />0.004 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (18 cm180 mm <br />0.18 m <br />) +
Contr. U.S. Natl. Herb. +
basal +  and cauline +
clavate +  and globose +
uniformly tan +  and brown +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
7 +  and numerous +
curved-cylindric +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
purple +, pink +, red +  and green +
deciduous +
0.55 cm5.5 mm <br />0.0055 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
unequal +
aquatic +, amphibious +  and terrestrial +
2 times as many or as many as sepals +
prostrate +  and decumbent +
globose +, lobed +  and entire +
intrapetiolar +
2 cm20 mm <br />0.02 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0.95 cm9.5 mm <br />0.0095 m <br /> (2 cm20 mm <br />0.02 m <br />) +
aquatic +, amphibious +  and terrestrial +
non-papillate +
Camissonia tanacetifolia +, C. tanacetifolia subsp. quadriperforata +, Taraxia longiflora +  and T. tikurana +
Taraxia tanacetifolia +
species +
3(-5)-aperturate +
distended +
smooth +  and erose +
papillate +