Thelypodium integrifolium

(Nuttall) Endlicher in W. G. Walpers

in W. G. Walpers, Repert. Bot. Syst. 1: 172. 1842.

Endemic
Basionym: Pachypodium integrifolium Nuttall in J. Torrey and A. Gray, Fl. N. Amer. 1: 96. 1838
Synonyms: Pleurophragma integrifolium (Nuttall) Rydberg
Treatment appears in FNA Volume 7. Treatment on page 732. Mentioned on page 728, 729.

Biennials; somewhat glaucous throughout, glabrous. Stems paniculately branched distally, (2–) 4.5–17 (–28) dm. Basal leaves: petiole (0.5–) 1.2–10 (–15) cm; blade usually oblong to lanceolate, or oblanceolate to spatulate, rarely ovate or obovate, (3.7–) 5–31 (–54) cm × (12–) 16–78 (–140) mm, margins usually entire, rarely dentate or repand. Cauline leaves (ascending, not appressed to stem); sessile; blade linear to linear-lanceolate, (1–) 2.2–8.3 (–19.5) cm × (2–) 3–11 (–25) mm, (base cuneate to attenuate, not auriculate), margins entire. Racemes dense, slightly or considerably elongated in fruit, (flower buds narrowly oblong). Fruiting pedicels usually horizontal to divaricate, rarely ascending or reflexed, usually straight, rarely incurved, slender or stout, (2–) 3–9 (–13) mm, often strongly flattened, rarely not flattened at base. Flowers: sepals erect, linear to linear-oblong, (3–) 3.5–5.5 (–7.5) × (0.5–) 0.8–1 (–1.5) mm; petals white or lavender to purple, spatulate to oblanceolate, (4.5–) 6–9 (–13) × (0.5–) 0.8–1.5 (–2.3) mm, margins not crisped, claw differentiated from blade, [slender, (2–) 3–4.5 (–7) mm, narrowest at base]; nectar glands lateral, flat or toothlike, median glands absent; filaments subequal to slightly tetradynamous, (3–) 4.5–7.5 (–12.5) mm; anthers exserted, linear to narrowly oblong, (1–) 1.5–2.5 (–4.5) mm, not circinately coiled; gynophore 0.5–2 (–5.5) mm. Fruits usually horizontal to divaricate-ascending, or ascending, rarely reflexed, usually torulose, rarely submoniliform, usually strongly incurved, sometimes straight, rarely arcuate, terete, (0.8–) 1.4–6.5 (–8) cm × 1–1.3 (–2) mm; ovules 14–40 per ovary; style usually cylindrical, rarely subclavate, 0.5–1.3 (–3.2) mm. Seeds (1–) 1.2–2 (–2.3) × 0.7–1 (–1.3) mm.

Distribution

V7 1278-distribution-map.gif

Ariz., Calif., Colo., Idaho, Mont., N.Dak., N.Mex., Nebr., Nev., Oreg., S.Dak., Utah, Wash., Wyo.

Discussion

Subspecies 5 (5 in the flora).

Selected References

None.

Key

1 Petals white; fruiting pedicels whitish, stout, (5-) 6-11(-13) mm > 2
1 Petals usually lavender to purple, rarely white; fruiting pedicels not whitish, slender and 2-10 (-13) mm, or stout and 2-5(-6) mm > 3
2 Petals (5-)6-9(-10) mm; fruiting pedicels horizontal to divaricate, (5-)6-9(-13) mm; fruits (1.4-)1.9-3.7(-4.5) cm; gynophores (0.5-)1-3 mm; se California, s Nevada, sw Utah. Thelypodium integrifolium subsp. affine
2 Petals (7.5-)8-11(-13) mm; fruiting pedicels usually horizontal to reflexed, rarely divaricate, (6-)7-11(-13) mm; fruits (2.2-) 3.5-6.5(-8) cm; gynophores (1-)1.5-4 (-5.5) mm; Grand Canyon, Colorado River, Arizona. Thelypodium integrifolium subsp. longicarpum
3 Fruiting pedicels often slender, sometimes stout, not or slightly flattened at base, (4-)6- 10 (-13) mm. Thelypodium integrifolium subsp. integrifolium
3 Fruiting pedicels stout, strongly flattened at base, 2-5(-6) mm > 4
4 Racemes often strongly congested, central rachises (0.6-)1.2-7(-9) cm; gynophores stout, 0.5-1(-3) mm; e California, Nevada, s Oregon, nw Utah. Thelypodium integrifolium subsp. complanatum
4 Racemes somewhat congested, central rachises (3-)8-20(-28) cm; gynophores slender, (0.5-)0.8-3(-4) mm; ne Arizona, w Colorado, nw New Mexico, e Utah. Thelypodium integrifolium subsp. gracilipes

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Thelypodium integrifolium"
0.25 cm2.5 mm <br />0.0025 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
linear +  and narrowly oblong +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
Ihsan A. Al-Shehbaz +
(Nuttall) Endlicher in W. G. Walpers +
not +  and rosulate +
straight +
incurved +, reflexed +, ascending +  and horizontal +
flattened +
stout +  and slender +
0.3 cm3 mm <br />0.003 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
Pachypodium integrifolium +
appendaged +  and unappendaged +
8.3 cm83 mm <br />0.083 m <br /> (19.5 cm195 mm <br />0.195 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
not +  and auriculate +
2.2 cm22 mm <br />0.022 m <br /> (8.3 cm83 mm <br />0.083 m <br />) +
laciniate +, lobed +, lyrate +  and dentate +
linear;linear-lanceolate +
much smaller +  and reduced +
0.3 cm3 mm <br />0.003 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
well-developed +
distinct +
differentiated +
emarginate +  and entire +
accumbent +  and incumbent +
Ariz. +, Calif. +, Colo. +, Idaho +, Mont. +, N.Dak. +, N.Mex. +, Nebr. +, Nev. +, Oreg. +, S.Dak. +, Utah +, Wash. +  and Wyo. +
straight +  and curved +
0.75 cm7.5 mm <br />0.0075 m <br /> (1.25 cm12.5 mm <br />0.0125 m <br />) +
connate +  and distinct +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
zygomorphic +  and actinomorphic +
purple;lavender;purple;lavender;white +
dilated +, spatulate +  and obovate or oblanceolate +
6.5 cm65 mm <br />0.065 m <br /> (8 cm80 mm <br />0.08 m <br />) +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
straight +
1.4 cm14 mm <br />0.014 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
incurved +, reflexed +, usually horizontal +  and divaricate-ascending or ascending +
terete +, submoniliform +  and torulose +
0.1 cm1 mm <br />0.001 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
not crisped;repand;dentate +
toothlike +  and flat +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
0.9 cm9 mm <br />0.009 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.23 cm2.3 mm <br />0.0023 m <br />) +
lavender +  and purple +
rudimentary +
spatulate +  and oblanceolate +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
10 cm100 mm <br />0.1 m <br /> (15 cm150 mm <br />0.15 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (10 cm100 mm <br />0.1 m <br />) +
trinucleate +  and 3(-11)-colpate +
in W. G. Walpers, Repert. Bot. Syst. +
reflexed +, erect +  and ascending +
stout +  and slender +
mucilaginous +
aseriate +, uniseriate +  and biseriate +
0.2 cm2 mm <br />0.002 m <br /> (0.23 cm2.3 mm <br />0.0023 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
ovate +, oblong +  and flattened +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
not basally +  and saccate +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
persistent +  and caducous +
distinct +
linear +  and linear-oblong +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
subequal +
procumbent +  and decumbent +
entire +  and capitate +
2-lobed +  and entire +
subsessile +, sessile +  and petiolate +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.32 cm3.2 mm <br />0.0032 m <br />) +
persistent +
distinct +
subclavate +  and cylindrical +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
Pleurophragma integrifolium +
Thelypodium integrifolium +
Thelypodium +
species +
dendritic +  and simple +
coiled +, 1-7-veined +  and veined +
anastomosing +
glabrous +  and glaucous +
perennial +, biennial +  and annual +
subshrub +  and shrub +
aquatics +  and terrestrial +
glabrous +  and pubescent +