Thelypodium wrightii

A. Gray

Smithsonian Contr. Knowl. 3(5): 7. 1852.

Synonyms: Stanleyella wrightii (A. Gray) Rydberg Thelypodium wrightii subsp. oklahomensis Al-Shehbaz
Treatment appears in FNA Volume 7. Treatment on page 738. Mentioned on page 729.

Biennials; slightly glaucous, glabrous. Stems branched distally, (6.5–) 9.5–22.5 (–28) dm. Basal leaves (and proximal cauline): petiole (2–) 4.5–8.5 (–13.7) cm; blade often lanceolate, sometimes oblanceolate in outline (lateral lobes oblong to linear or deltate), (6.5–) 9.5–22.5 (–28) cm × 28–55 (–75) mm, margins pinnately lobed or lyrate (lobes entire or denticulate). Cauline leaves petiolate; blade lanceolate to linear-lanceolate, 3.5–8.5 (–13.5) cm × 7–14 (–23) mm, margins usually entire or dentate, rarely lobed. Racemes somewhat lax, considerably elongated in fruit. Fruiting pedicels horizontal to divaricate, sometimes reflexed, straight or curved, sometimes secund, slender, (5–) 7–13 (–17) mm, slightly flattened at base. Flowers: sepals spreading or ascending oblong to linear, (3–) 4–6 (–7) × (0.8–) 1–1.3 (–1.5) mm; petals usually white, rarely lavender, linear or oblong, 4–7.5 (–9) × 1–1.8 (–2) mm, margins not crisped, claw not differentiated from blade (base sometimes clawlike, to 2 mm, widest at base); nectar glands confluent; filaments equal, (3–) 3.5–6.5 (–8.5) mm; anthers (1–) 1.5–2.5 (–3) mm, circinately coiled; gynophore 0.2–2 (–5) mm. Fruits horizontal or reflexed, torulose, straight to somewhat curved, flattened, (2.5–) 3.8–7.4 (–9) cm × 1–1.2 (–1.5) mm, (replum not constricted between seeds); ovules 76–128 per ovary; style usually clavate to subclavate, rarely cylindrical, (0.5–) 0.8–2 (–3) mm. Seeds 0.7–1.3 (–1.5) × 0.5–0.8 mm.


Phenology: Flowering (Mar-)Jun–Oct.
Habitat: Rock crevices, shady slopes, rocky hillsides, canyons, stream banks, creek beds, pinyon-juniper communities, oak woodlands
Elevation: 1200-2300 m

Distribution

V7 1295-distribution-map.gif

Ariz., Colo., N.Mex., Tex., Utah, Mexico (Baja California), Mexico (Chihuahua), Mexico (Coahuila), Mexico (Hidalgo)

Discussion

Study of extensive material that has accumulated during the past three decades reveals that the differences between subsp. oklahomensis and subsp. wrightii, which are based solely on variations in gynophore and fruit lengths, do not hold as clearly as previously thought (I. A. Al-Shehbaz 1973; R. C. Rollins 1993).

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Thelypodium wrightii"
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
usually linear +  and linear-oblong rarely oblong or ovate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
Ihsan A. Al-Shehbaz +
A. Gray +
not +  and rosulate +
curved +  and straight +
reflexed +  and horizontal +
flattened +
slender +
0.7 cm7 mm <br />0.007 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
Pachypodium +
appendaged +  and unappendaged +
8.5 cm85 mm <br />0.085 m <br /> (13.5 cm135 mm <br />0.135 m <br />) +
1.4 cm14 mm <br />0.014 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
not +  and auriculate +
3.5 cm35 mm <br />0.035 m <br /> (8.5 cm85 mm <br />0.085 m <br />) +
laciniate +, lobed +, lyrate +  and dentate +
lanceolate;linear-lanceolate +
much smaller +  and reduced +
0.7 cm7 mm <br />0.007 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
well-developed +
distinct +
not differentiated +
emarginate +  and entire +
accumbent +  and incumbent +
Ariz. +, Colo. +, N.Mex. +, Tex. +, Utah +, Mexico (Baja California) +, Mexico (Chihuahua) +, Mexico (Coahuila) +  and Mexico (Hidalgo) +
1200-2300 m +
straight +  and curved +
not winged +  and unappendaged +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
connate +  and distinct +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
zygomorphic +  and actinomorphic +
purple;lavender;purple;lavender;white +
dilated +, spatulate +  and obovate or oblanceolate +
7.4 cm74 mm <br />0.074 m <br /> (9 cm90 mm <br />0.09 m <br />) +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
curved +  and straight to somewhat +
3.8 cm38 mm <br />0.038 m <br /> (7.4 cm74 mm <br />0.074 m <br />) +
reflexed +  and horizontal +
flattened +  and torulose +
0.1 cm1 mm <br />0.001 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Rock crevices, shady slopes, rocky hillsides, canyons, stream banks, creek beds, pinyon-juniper communities, oak woodlands +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
dentate +  and entire +
not crisped;lobed;lyrate;lobed +
22.5 cm225 mm <br />0.225 m <br /> (28 cm280 mm <br />0.28 m <br />) +
5.5 cm55 mm <br />0.055 m <br /> (7.5 cm75 mm <br />0.075 m <br />) +
9.5 cm95 mm <br />0.095 m <br /> (22.5 cm225 mm <br />0.225 m <br />) +
2.8 cm28 mm <br />0.028 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
0.75 cm7.5 mm <br />0.0075 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
4mm;7.5mm +
rudimentary +
oblong +  and linear +
0.1 cm1 mm <br />0.001 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
8.5 cm85 mm <br />0.085 m <br /> (13.7 cm137 mm <br />0.137 m <br />) +
4.5 cm45 mm <br />0.045 m <br /> (8.5 cm85 mm <br />0.085 m <br />) +
Flowering (Mar-)Jun–Oct. +
trinucleate +  and 3(-11)-colpate +
Smithsonian Contr. Knowl. +
reflexed +, erect +  and ascending +
stout +  and slender +
mucilaginous +
aseriate +, uniseriate +  and biseriate +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
ovate +, oblong +  and flattened +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
not basally +  and saccate +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
persistent +  and caducous +
distinct +
ascending oblong +  and linear +
ovate +  and oblong linear lanceolate or oblanceolate +
0.1 cm1 mm <br />0.001 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
subequal +
procumbent +  and decumbent +
entire +  and capitate +
2-lobed +  and entire +
subsessile +, sessile +  and petiolate +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
persistent +
distinct +
cylindrical +, usually clavate +  and subclavate +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Stanleyella wrightii +  and Thelypodium wrightii subsp. oklahomensis +
Thelypodium wrightii +
Thelypodium +
species +
dendritic +  and simple +
coiled +, 1-7-veined +  and veined +
anastomosing +
glabrous +  and glaucous +
perennial +, biennial +  and annual +
subshrub +  and shrub +
aquatics +  and terrestrial +
glabrous +  and pubescent +