Timmiella anomala

(Bruch & Schimper) Limpricht

Laubm. Deutschl. 1: 592. 1888,.

Basionym: Barbula anomala Bruch & Schimper Bryol. Europ. 2: 107, plate 169. 1842
Synonyms: Timmiella subanomala (Bescherelle) Brotherus Trichostomum subanomala (Bescherelle) Brotherus
Treatment appears in FNA Volume 27. Treatment on page 482.

Stems commonly branched. Leaf base narrower to little wider than the limb. Sexual condition dioicous, autoicous or synoicous, occasionally polyoicous. Seta usually 1.5–2.5 cm. Capsule with theca reaching 3–4 mm, operculum commonly 1–1.5 mm, peristome distinctly twisted.


Phenology: Sporophytes mature summer–fall (occasionally winter).
Habitat: Soil, humus, limestone crevices, vertical rock face
Elevation: moderate elevations (700-1500 m)

Distribution

V27 697-distribution-map.gif

Ariz., Calif., N.Mex., Tex., Wash., Mexico, Central America, Europe, Asia, n Africa

Discussion

Timmiella anomala commonly grows in dusty, friable soil that makes dissections difficult. H. A. Crum and L. E. Anderson (1980–1983, vol. 4) suggested that this species may also be dioicous; K. Saito (1975) and Li X. J. et al. (2001) asserted that it is dioicous in Asia (Japan and China, respectively). A specimen from California, Showers 3326, MO, has stems with synoicous perichaetia and also apparently entirely perigoniate stems, possibly due to rhizautoicy. According to B. Allen (2002), Timmiella anomala in Central America has peristome teeth erect to very weakly twisted though these are described as at least once twisted in publications on other areas of the world. Material from the Yukon is sterile and only tentatively identifiable as this species.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property.

... more about "Timmiella anomala"
3-stratose +  and 2-stratose +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
distinct +
of 1;several layers +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
Richard H. Zander +
(Bruch & Schimper) Limpricht +
crescent--shaped +
papillose +  and smooth +
usually slightly wider +
usually ovate;oblong +
broadened +
Barbula anomala +
obovoid +
cucullate +
stegocarpous +
monoicous +, dioicous +  and sexual +
flat to bulging +
vesiculose +  and 2-stratose +
specialized +
monoicous +, dioicous +  and not asexual +
elongate-rhomboid +  and elongate +
tomentose +  and bare +
tapering +
broadened +
serrate +  and serrulate +
denticulate +  and plane +
quadrate;rounded-hexagonal +
Ariz. +, Calif. +, N.Mex. +, Tex. +, Wash. +, Mexico +, Central America +, Europe +, Asia +  and n Africa +
moderate elevations (700-1500 m) +
differentiated +
multicellular +
Soil, humus, limestone crevices, vertical rock face +
yellow +  and orange-red or red +
1-stratose +  and 2-stratose +
1 +  and several +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
polyoicous +, synoicous +, autoicous +, dioicous +  and sexual +
wider +  and narrower +
lanceolate +
reduced +
rounded +
dentate +  and entire +
long-conic;rostrate +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
photosynthetic +
compound +, simple +, hollow +  and solid +
flattened +  and 2-fid +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
Sporophytes mature summer–fall (occasionally winter). +
1 (?) +, 3 (?) +  and 4 (?) +
Laubm. Deutschl. +
straight +
elongate +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
2 +  and 1 +
yellow +  and laminal +
rounded-triangular +  and rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
Timmiella subanomala +  and Trichostomum subanomala +
Timmiella anomala +
Timmiella +
species +
ventricose;long-elliptic;cylindric +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
straight +
hyaline +  and yellowish +
wasp-waisted +  and lanceolate +
stemless +  and perigoniate +
plant +, loosely cespitose +  and cushion-forming +
occasionally smaller than the perichaetiate +