Tortula bolanderi

(Lesquereu×& James) M. Howe

Erythea 4: 51. 1896,.

Basionyms: Barbula bolanderi Lesquereux & James Trans. Amer. Philos. Soc., n. s. 13: 5. 1896
Synonyms: Barbula carnifolia Müller Hal. & Kindberg Syntrichia bolanderi (Lesquereux) R. H. Zander
Treatment appears in FNA Volume 27. Treatment on page 602. Mentioned on page 587, 588, 590, 603.

Leaves long-ovate, elliptic or lingulate, apex rounded and short-apiculate or muticuous, margins weakly recurved in proximal 1/2 of leaf, weakly bordered in distal 1/2–3/4 of leaf with (2–) 4–6 rows of thicker-walled, smaller cells; costa subpercurrent, percurrent or shortly excurrent as a small, sharp mucro, lacking an adaxial pad of cells, distally somewhat narrow, 3–4 cells across adaxial surface; distal laminal cells hexagonal, width 13–18 (–24) µm, 1: 1, strongly papillose. Sexual condition dioicous. Sporophytes exerted. Seta 0.9–1.5 cm. Capsule stegocarpic, not systylius, cylindric, erect and nearly straight, urn 1.5–2.5 mm; peristome of 32 twisted filaments, basal membrane low; operculum 1–1.5 mm. Spores 13–15 µm, spheric, essentially smooth.


Habitat: Soil, rock
Elevation: low to high elevations (0-1700 m)

Distribution

V27 868-distribution-map.gif

B.C., Calif., Oreg., Wash., Europe (France), Europe (Spain)

Discussion

Both Tortula bolanderi and T. amplexa differ from other species of the genus in the red color of the leaves in nature, a trait of Syntrichia. The costal section, however, exhibits a rounded stereid band rather than the semilunate shape of the latter.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property.

... more about "Tortula bolanderi"
smooth +  and papillose +
smooth +  and papillose +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
channeled +, nearly flat +  and concave +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
Richard H. Zander +  and Patricia M. Eckel +
(Lesquereu×& James) M. Howe +
semicircular +  and elliptical +
semicircular +, elliptical +  and rounded +
papillose +  and smooth +
usually slightly wider +
usually ovate;oblong +
differentiated +
Barbula bolanderi +
obovoid +
cucullate +
0.25 cm2.5 mm <br />0.0025 m <br /> (?) +  and 0.6 cm6 mm <br />0.006 m <br /> (?) +
not systylius +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
dark-brown +  and yellowish-brown +
cleistocarpous +  and stegocarpous +
cylindric +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
monoicous +, dioicous +  and sexual +
twisted +, vesiculose +  and thicker-walled +
1 (?) +  and 3 (?) +
elongate-rhomboid +  and elongate +
tomentose +  and bare +
excurrent +  and percurrent +
subpercurrent +
dioicous +  and sexual +
B.C. +, Calif. +, Oreg. +, Wash. +, Europe (France) +  and Europe (Spain) +
low to high elevations (0-1700 m) +
differentiated +
multicellular +
Soil, rock +
somewhat larger +
yellow +  and orange-red or red +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
lingulate +, elliptic +  and long-ovate +
reduced +
serrulate +  and entire +
smaller +
rostrate;long-conic +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
differentiated +
photosynthetic +
simple +  and hollow +
4 +  and 6 +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
straight +
not twisted +  and twisted +
brown;yellowish-brown +
elongate +
0.9 cm9 mm <br />0.009 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
2 +  and 1 +
orange +, red +, yellow +, laminal +  and light-brown +
spheric +
rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Barbula carnifolia +  and Syntrichia bolanderi +
Tortula bolanderi +
species +
ovoid;cylindric +
rudimentary +
32 +  and 16 +
triangular +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
yellow distally and red +, orange +, red +  and yellow +
rectangular +
thickened +, thin +  and thicker +
stemless +  and perigoniate +
yellowbrown;dark-brown +
occasionally smaller than the perichaetiate +