Tortula brevipes

(Lesquereux) Brotherus in H. G. A. Engler and K. Prantl

in H. G. A. Engler and K. Prantl, Nat. Pflanzenfam. 214[I,3]: 431. 1902,.

Basionyms: Barbula brevipes Lesquereux Mem. Calif. Acad. Sci. 1: 12. 1868
Synonyms: Barbula macrotrichia Cardot & Thériot
Treatment appears in FNA Volume 27. Treatment on page 595. Mentioned on page 589, 590, 596.

Leaves short-elliptic or long-obovate, sometimes ovate, apex very broadly acute to rounded, short to long-awned, margins very broadly recurved or revolute to near apex, not bordered; costa usually long-excurrent, lacking an adaxial pad of cells, distally narrow, ca. 3–4 cells across the convex adaxial surface; distal laminal cells hexagonal, 10–15 µm wide, 1: 1, densely papillose with 2-fid papillae. Sexual condition apparently dioicous but more probably rhizautoicous, occasionally autoicous. Sporophytes exerted. Seta length ca. 0.7–1.4 (–2) cm. Capsule stegocarpic, not systylius, cylindric, erect and nearly straight, urn length ca. (1.7–) 2.5–3.5 mm; peristome 800–1000 µm, teeth of 32 filaments twisted about one full turn, basal membrane 150–200 µm; operculum 1–1.5 mm. Spores ca. 10–13 µm, spheric, finely papillose.


Habitat: Calcareous soil
Elevation: low to moderate elevations

Distribution

V27 850-distribution-map.gif

Ariz., Calif., Colo., Idaho, Nev., Oreg., Utah, Wash., Mexico (Baja California), Mexico (Sonora)

Discussion

Tortula brevipes is much like T. muralis, but differs in the soil substrate, absence of a leaf border (common in T. muralis), the apparently dioicous inflorescence, the commonly very long (to 5 mm) operculate capsule, and the well-developed basal membrane of the peristome. According to B. D. Mishler (1994), T. brevipes in Mexico intergrades with T. muralis in capsule length, height of the basal membrane, and development of a leaf border.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property."um" is not declared as a valid unit of measurement for this property.

... more about "Tortula brevipes"
smooth +  and papillose +
smooth +  and papillose +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
Richard H. Zander +  and Patricia M. Eckel +
(Lesquereux) Brotherus in H. G. A. Engler and K. Prantl +
semicircular +  and elliptical +
semicircular +, elliptical +  and rounded +
papillose +  and smooth +
usually slightly wider +
usually ovate;oblong +
differentiated +
Barbula brevipes +
obovoid +
cucullate +
0.25 cm2.5 mm <br />0.0025 m <br /> (?) +  and 0.6 cm6 mm <br />0.006 m <br /> (?) +
not systylius +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
dark-brown +  and yellowish-brown +
cleistocarpous +  and stegocarpous +
cylindric +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
monoicous +, dioicous +  and sexual +
twisted +, vesiculose +  and thicker-walled +
1 (?) +  and 3 (?) +
elongate-rhomboid +  and elongate +
tomentose +  and bare +
long-excurrent +
subpercurrent +
autoicous +, dioicous +  and sexual +
Ariz. +, Calif. +, Colo. +, Idaho +, Nev. +, Oreg. +, Utah +, Wash. +, Mexico (Baja California) +  and Mexico (Sonora) +
low to moderate elevations +
differentiated +
multicellular +
Calcareous soil +
somewhat larger +
yellow +  and orange-red or red +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
ovate +, long-obovate +  and short-elliptic +
reduced +
serrulate +  and entire +
smaller +
rostrate;long-conic +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
differentiated +
photosynthetic +
simple +  and hollow +
4 +  and 6 +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
in H. G. A. Engler and K. Prantl, Nat. Pflanzenfam. +
straight +
not twisted +  and twisted +
1.4 cm14 mm <br />0.014 m <br /> (2 cm20 mm <br />0.02 m <br />) +
brown;yellowish-brown +
0.7 cm7 mm <br />0.007 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
elongate +
0 cm0 mm <br />0 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
2 +  and 1 +
orange +, red +, yellow +, laminal +  and light-brown +
spheric +
rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Barbula macrotrichia +
Tortula brevipes +
species +
ovoid;cylindric +
rudimentary +
32 +  and 16 +
triangular +
0.17 cm1.7 mm <br />0.0017 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
yellow distally and red +, orange +, red +  and yellow +
rectangular +
thickened +, thin +  and thicker +
stemless +  and perigoniate +
yellowbrown;dark-brown +
occasionally smaller than the perichaetiate +