Tortula californica

E. B. Bartram

Bryologist 48: 92, figs. h–n. 1945,.

Treatment appears in FNA Volume 27. Treatment on page 593. Mentioned on page 588, 590, 594.

Leaves broadly obovate, occasionally elliptic, apex broadly acute, rounded or occasionally emarginate, awned, margins plane or occasionally weakly recurved proximally, often weakly bordered distally with slightly smaller and slightly more thick-walled cells; costa long-excurrent, lacking an adaxial pad of cells, strong but distally narrow, mostly 2–4 cells across adaxial surface; distal laminal cells inflated hexagonal, occasionally irregularly quadrate, width 18–25 µm, 1: 1 or occasionally longitudinally elongate, smooth. Sexual condition autoicous. Sporophytes exerted. Seta 0.9–1.1 cm. Capsule stegocarpic, not systylius, cylindric, erect and nearly straight, urn 2–2.7 mm; peristome 1000–1400 µm, teeth of 32 filaments, twisted once, basal membrane ca. 200 µm; operculum 1.1–1.5 mm. Spores 11–14 µm, spheric, essentially smooth.


Phenology: Capsules mature spring.
Habitat: Arid soil, rock
Elevation: low elevations (0-50 m)

Distribution

V27 847-distribution-map.gif

Calif., Mexico (Baja California)

Discussion

Tortula californica has now been reported from several counties in southern California (D. H. Norris and J. R. Shevock 2004), and is characterized by obovate leaves with smooth awns, plane margins, and smooth laminal cells.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property."um" is not declared as a valid unit of measurement for this property.

... more about "Tortula californica"
smooth +  and papillose +
smooth +  and papillose +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
channeled +, nearly flat +  and concave +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
awned +  and emarginate +
Rounded (?) +  and Acute (?) +
Richard H. Zander +  and Patricia M. Eckel +
E. B. Bartram +
semicircular +  and elliptical +
semicircular +, elliptical +  and rounded +
papillose +  and smooth +
usually slightly wider +
usually ovate;oblong +
differentiated +
obovoid +
cucullate +
0.25 cm2.5 mm <br />0.0025 m <br /> (?) +  and 0.6 cm6 mm <br />0.006 m <br /> (?) +
not systylius +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
dark-brown +  and yellowish-brown +
cleistocarpous +  and stegocarpous +
cylindric +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
monoicous +, dioicous +  and sexual +
thick-walled +
1 (?) +  and 3 (?) +
elongate-rhomboid +  and elongate +
smaller +
tomentose +  and bare +
long-excurrent +
subpercurrent +
autoicous +  and sexual +
Calif. +  and Mexico (Baja California) +
low elevations (0-50 m) +
differentiated +
multicellular +
Arid soil, rock +
somewhat larger +
yellow +  and orange-red or red +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
obovate +
reduced +
serrulate +  and entire +
smaller +
rostrate;long-conic +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
differentiated +
photosynthetic +
simple +  and hollow +
4 +  and 6 +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
Capsules mature spring. +
straight +
not twisted +  and twisted +
brown;yellowish-brown +
elongate +
0.9 cm9 mm <br />0.009 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
2 +  and 1 +
orange +, red +, yellow +, laminal +  and light-brown +
spheric +
rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Desmatodon +, Phascum +, Pottia +  and Protobryum +
Tortula californica +
species +
ovoid;cylindric +
rudimentary +
32 +  and 16 +
triangular +
0.2 cm2 mm <br />0.002 m <br /> (0.27 cm2.7 mm <br />0.0027 m <br />) +
yellow distally and red +, orange +, red +  and yellow +
rectangular +
thickened +, thin +  and thicker +
stemless +  and perigoniate +
yellowbrown;dark-brown +
occasionally smaller than the perichaetiate +