Tortula cernua

(Huebener) Lindberg

Musc. Scand., 20. 1879,.

Basionyms: Dermatodon cernuus Huebener Muscol. Germ., 117. 1833
Synonyms: Dermatodon randii (Kennedy) Lazarenko Pottia randii Kennedy Tortula randii (Kennedy) R. H. Zander
Treatment appears in FNA Volume 27. Treatment on page 601. Mentioned on page 587, 589, 590, 593, 599, 602.

Leaves elliptic, apex broadly acute to rounded-acute, mucronate or apiculate, margins recurved at mid leaf, bordered throughout or in lower 2/3 with 2 (–3) rows of narrower, short to long-rectangular, thicker-walled cells that 1–2-stratose; costa percurrent or short-excurrent, lacking an adaxial pad of cells, distally narrow, 3–4 cells across the convex adaxial surface; distal laminal cells hexagonal to rectangular, (13–) 16–24 (–28) µm wide, 1–2: 1, not or rarely very weakly papillose. Sexual condition autoicous. Sporophytes exerted. Seta 0.6–1.2 cm. Capsule stegocarpic, not systylius, short-cylindric and curved or ovoid and bilaterally symmetrical, nodding or occasionally nearly horizontal, urn 1–1.8 mm; peristome weak, ca. 100–200 µm, teeth 16, not twisted, divided nearly to the base into 2–3 branches and variously perforate, basal membrane very low or absent, or rudimentary or fragmentary and attached to the inside of the operculum; operculum 0.2–1.5 mm. Spores 25–35 µm, spheric, densely papillose.


Phenology: Capsules mature summer.
Habitat: Soil, limestone
Elevation: low to moderate elevations

Distribution

V27 865-distribution-map.gif

Greenland, Alta., B.C., Man., N.B., Nfld. and Labr. (Nfld.), N.W.T., N.S., Nunavut, Ont., Que., Sask., Yukon, Alaska, Ariz., Colo., Ill., Iowa, Maine, Mont., Nev., N.Dak., Ohio, Utah, Wis., Wyo., Europe, Asia

Discussion

Tortula cernua differs from T. laureri in the usually smooth laminal cells, the mostly ovoid and bilaterally symmetrical capsule inclined to horizontal, and peristome not twisted. Its range is more southerly and at lower elevations. The small and fragile peristome commonly falls with the operculum, being attached to its inner surface, and the capsule may appear gymnostomous. Given that the uncommon T. randii is only distinguished from T. cernua by the apparent absence of a peristome and has much the same range, and all specimens of that species I have seen are indeed peristomate or, as in A. J. Grout, N. Amer. Musci Perf. 197, MO, have smooth fragments of a peristome attached to the inner surface of the operculum, I agree with J. M. Holzinger (1925) that it should not be recognized as distinct.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property."um" is not declared as a valid unit of measurement for this property.

... more about "Tortula cernua"
smooth +  and papillose +
smooth +  and papillose +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
Richard H. Zander +  and Patricia M. Eckel +
(Huebener) Lindberg +
semicircular +  and elliptical +
semicircular +, elliptical +  and rounded +
papillose +  and smooth +
usually slightly wider +
fragmentary +  and rudimentary +
usually ovate;oblong +
differentiated +
Dermatodon cernuus +
obovoid +
cucullate +
0.25 cm2.5 mm <br />0.0025 m <br /> (?) +  and 0.6 cm6 mm <br />0.006 m <br /> (?) +
not systylius +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
dark-brown +  and yellowish-brown +
cleistocarpous +  and stegocarpous +
horizontal +  and nodding +
ovoid;short-cylindric +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
monoicous +, dioicous +  and sexual +
thicker-walled +
1 (?) +  and 3 (?) +
long-rectangular +
narrower +
tomentose +  and bare +
short-excurrent +  and percurrent +
subpercurrent +
autoicous +  and sexual +
Greenland +, Alta. +, B.C. +, Man. +, N.B. +, Nfld. and Labr. (Nfld.) +, N.W.T. +, N.S. +, Nunavut +, Ont. +, Que. +, Sask. +, Yukon +, Alaska +, Ariz. +, Colo. +, Ill. +, Iowa +, Maine +, Mont. +, Nev. +, N.Dak. +, Ohio +, Utah +, Wis. +, Wyo. +, Europe +  and Asia +
low to moderate elevations +
differentiated +
multicellular +
Soil, limestone +
somewhat larger +
yellow +  and orange-red or red +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
obovate +  and spathulate ovate +
reduced +
in lower 2/3 +  and bordered +
serrulate +  and entire +
smaller +
rostrate;long-conic +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
differentiated +
photosynthetic +
simple +  and hollow +
4 +  and 6 +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
Capsules mature summer. +
Musc. Scand., +
1-2-stratose +
straight +
not twisted +  and twisted +
brown;yellowish-brown +
elongate +
0.6 cm6 mm <br />0.006 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
2 +  and 1 +
orange +, red +, yellow +, laminal +  and light-brown +
spheric +
rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Dermatodon randii +, Pottia randii +  and Tortula randii +
Tortula cernua +
species +
ovoid;cylindric +
perforate +  and not twisted +
rudimentary +
divided +
0.1 cm1 mm <br />0.001 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
yellow distally and red +, orange +, red +  and yellow +
rectangular +
thickened +, thin +  and thicker +
stemless +  and perigoniate +
yellowbrown;dark-brown +
occasionally smaller than the perichaetiate +