Tortula hoppeana

(Schultz) Ochyra

Bryologist 107: 499. 2005,.

Basionyms: Trichostomum hoppeanum Schultz in C. F. Hornschuch, Syll. Pl. Nov. 2: 140. 1828
Synonyms: Desmatodon latifolius (Hedwig) Bridel Desmatodon latifolius var. muticus (Bridel) Bridel Tortula eucalyptrata R. H. Zander Tortula euryphylla
Treatment appears in FNA Volume 27. Treatment on page 598. Mentioned on page 588, 589, 590, 599, 600.

Leaves ovate to oblong, apex broadly acute or occasionally rounded, apiculate or short to long-mucronate, occasionally short-awned, margins recurved in middle 2/3 of leaf or occasionally almost plane, not or very weakly bordered distally with ca. 2 rows of rhomboid, less papillose cells; costa subpercurrent, percurrent or excurrent, lacking an adaxial pad of cells, distally narrow, 2–3 cells across the convex adaxial surface; distal laminal cells hexagonal, 15–20 µm wide, 1: 1, densely papillose with several 2-fid papillae. Sexual condition autoicous. Sporophytes exerted. Seta 0.9–1.2 (–1.9) cm. Capsule stegocarpic, rarely systylius, cylindric, erect and nearly straight, urn 1.5–2 (–2.8) mm; peristome (150–) 300–350 length µm, teeth 16, usually not twisted, divided to near the base or perforate basally, the teeth often basally fused in 2s or 4s, basal membrane very low; operculum 0.7–1 mm. Spores ca. 20–23 (–25) µm, spheric or elliptic, densely papillose.


Phenology: Capsules mature summer.
Habitat: Soil, calcareous silt
Elevation: low (in northern regions) to high elevations of over 3300 m

Distribution

V27 857-distribution-map.gif

Greenland, Alta., B.C., Nfld. and Labr., N.W.T., Nunavut, Que., Yukon, Alaska, Ariz., Calif., Colo., Idaho, Mich., Mont., Nev., N.Mex., Oreg., Utah, Wash., Wyo., Europe, Asia, n Africa, Atlantic Islands (Iceland)

Discussion

Care must be taken to not be confused by recent nomenclatural changes involving the epithet “latifolia” in the Pottioideae. Pottia latifolia is now to be found in Stegonia; Desmatodon latifolius is now Tortula hoppeana; Tortula latifolia is now Syntrichia latifolia. The strong 2-fid laminal papillae of Tortula hoppeana are distinctive, contributing to a crenulate leaf margin at the apex, which is surmounted, excepting specimens with muticous leaves, by a narrow, long mucro of smooth, clear, elongate cells. This is a very common species in subalpine and alpine areas, and in tundra. The present treatment agrees with modern perception (L. E. Anderson et al. 1990; M. J. Cano et al. 2006) that the muticous-leaved variant, which has much the same distribution as the piliferous expression, is not worth recognizing. The peristome teeth are essentially 16 but are commonly paired or in larger groups. Like T. cernua, T. laureri, T. leucostoma, T. obtusifolia, and T. systylia, this species has a capsule mainly maturing in summer, a feature supporting recognition of the group under the name Desmatodon, but possibly at only the subgenus level.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property.

... more about "Tortula hoppeana"
smooth +  and papillose +
smooth +  and papillose +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
short-awned +  and apiculate +
Richard H. Zander +  and Patricia M. Eckel +
(Schultz) Ochyra +
semicircular +  and elliptical +
semicircular +, elliptical +  and rounded +
papillose +  and smooth +
usually slightly wider +
usually ovate;oblong +
differentiated +
Trichostomum hoppeanum +
obovoid +
cucullate +
0.25 cm2.5 mm <br />0.0025 m <br /> (?) +  and 0.6 cm6 mm <br />0.006 m <br /> (?) +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
dark-brown +  and yellowish-brown +
cleistocarpous +  and stegocarpous +
cylindric +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
monoicous +, dioicous +  and sexual +
twisted +, vesiculose +  and thicker-walled +
1 (?) +  and 3 (?) +
papillose +
rhomboid +
tomentose +  and bare +
excurrent +  and percurrent +
subpercurrent +
autoicous +  and sexual +
Greenland +, Alta. +, B.C. +, Nfld. and Labr. +, N.W.T. +, Nunavut +, Que. +, Yukon +, Alaska +, Ariz. +, Calif. +, Colo. +, Idaho +, Mich. +, Mont. +, Nev. +, N.Mex. +, Oreg. +, Utah +, Wash. +, Wyo. +, Europe +, Asia +, n Africa +  and Atlantic Islands (Iceland) +
low (in northern regions) to high elevations of over 3300 m +
differentiated +
multicellular +
Soil, calcareous silt +
somewhat larger +
yellow +  and orange-red or red +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
ovate +  and oblong +
reduced +
serrulate +  and entire +
smaller +
rostrate;long-conic +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
differentiated +
photosynthetic +
simple +  and hollow +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
Capsules mature summer. +
straight +
not twisted +  and twisted +
1.2 cm12 mm <br />0.012 m <br /> (1.9 cm19 mm <br />0.019 m <br />) +
brown;yellowish-brown +
elongate +
0.9 cm9 mm <br />0.009 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
2 +  and 1 +
orange +, red +, yellow +, laminal +  and light-brown +
elliptic +  and spheric +
rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Desmatodon latifolius +, Desmatodon latifolius var. muticus +, Tortula eucalyptrata +  and Tortula euryphylla +
Tortula hoppeana +
species +
ovoid;cylindric +
perforate +  and twisted +
rudimentary +
divided +
0.2 cm2 mm <br />0.002 m <br /> (0.28 cm2.8 mm <br />0.0028 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
yellow distally and red +, orange +, red +  and yellow +
rectangular +
thickened +, thin +  and thicker +
stemless +  and perigoniate +
yellowbrown;dark-brown +
occasionally smaller than the perichaetiate +