Tortula inermis

(Bridel) Montagne

Arch. Bot. (Leipzig) 1: 136. 1832,.

Basionyms: Syntrichia subulata var. inermis Bridel Bryol. Univ. 1: 581. 1826
Synonyms: Syntrichia inermis
Treatment appears in FNA Volume 27. Treatment on page 600. Mentioned on page 588, 590, 602.

Leaves lingulate, apex rounded-acute, apiculate to short-mucronate, margins recurved from base to near the apex, not bordered; costa subpercurrent, percurrent or very short-excurrent, lacking an adaxial pad of cells but distally narrowing or weakly thickened, 3–4 (–5) cells across the convex adaxial surface; distal laminal cells hexagonal, (12–) 15–18 µm wide, 1: 1, strongly papillose with 4–5 2-fid papillae. Sexual condition gonioautoicous. Sporophytes exerted. Seta 1.2–1.5 (–2.5) cm. Capsule stegocarpic, not systylius, cylindric, erect and nearly straight, urn usually 3–4 mm; peristome length 1000–1500 µm, teeth of 32 filaments twisted at least one full turn, basal membrane 300–600 µm; operculum 1.3–1.6 (–2) mm. Spores 11–15 µm, spheric, finely papillose.


Phenology: Capsules mature spring.
Habitat: Soil, rock
Elevation: low to high elevations (0-2000 m)

Distribution

V27 861-distribution-map.gif

Ariz., Calif., Colo., Nev., N.Mex., S.Dak., Tex., Utah, Wyo., Mexico (Baja California), Mexico (Chihuahua), Mexico (Sonora), Europe, s Asia, n Africa

Discussion

Tortula inermis has the aspect of a Syntrichia with its ligulate, apiculate leaves and strong costa, but the plant is yellow or orange in KOH solution, and the costal section reveals a rounded stereid band. It is related to T. subulata and T. mucronifolia but the lack of a strong mucro and the narrow but nearly complete recurving of the leaf margins are diagnostic.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property.

... more about "Tortula inermis"
smooth +  and papillose +
smooth +  and papillose +
thickened +  and narrowing +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
Richard H. Zander +  and Patricia M. Eckel +
(Bridel) Montagne +
semicircular +  and elliptical +
semicircular +, elliptical +  and rounded +
papillose +  and smooth +
usually slightly wider +
usually ovate;oblong +
differentiated +
Syntrichia subulata var. inermis +
obovoid +
cucullate +
0.25 cm2.5 mm <br />0.0025 m <br /> (?) +  and 0.6 cm6 mm <br />0.006 m <br /> (?) +
not systylius +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
dark-brown +  and yellowish-brown +
cleistocarpous +  and stegocarpous +
cylindric +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
monoicous +, dioicous +  and sexual +
twisted +, vesiculose +  and thicker-walled +
1 (?) +  and 3 (?) +
elongate-rhomboid +  and elongate +
tomentose +  and bare +
short-excurrent +  and percurrent +
subpercurrent +
Ariz. +, Calif. +, Colo. +, Nev. +, N.Mex. +, S.Dak. +, Tex. +, Utah +, Wyo. +, Mexico (Baja California) +, Mexico (Chihuahua) +, Mexico (Sonora) +, Europe +, s Asia +  and n Africa +
low to high elevations (0-2000 m) +
differentiated +
multicellular +
Soil, rock +
somewhat larger +
yellow +  and orange-red or red +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
lingulate +
reduced +
serrulate +  and entire +
smaller +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
rostrate;long-conic +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
differentiated +
photosynthetic +
simple +  and hollow +
4 +  and 6 +
4-5-2-fid +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
Capsules mature spring. +
Arch. Bot. (Leipzig) +
straight +
not twisted +  and twisted +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
brown;yellowish-brown +
elongate +
1.2 cm12 mm <br />0.012 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
2 +  and 1 +
orange +, red +, yellow +, laminal +  and light-brown +
spheric +
rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Syntrichia inermis +
Tortula inermis +
species +
ovoid;cylindric +
rudimentary +
32 +  and 16 +
triangular +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
yellow distally and red +, orange +, red +  and yellow +
rectangular +
thickened +, thin +  and thicker +
stemless +  and perigoniate +
yellowbrown;dark-brown +
occasionally smaller than the perichaetiate +