Tortula lanceola

R. H. Zander

Bull. Buffalo Soc. Nat. Sci. 32: 223. 1993,.

Basionyms: Encalypta lanceolata Hedwig Sp. Musc. Frond., 63. 1801,
Synonyms: Pottia lanceolata (Hedwig) Müller Hal.
Treatment appears in FNA Volume 27. Treatment on page 592. Mentioned on page 589.

Leaves obovate, elliptic or less commonly ovate, apex broadly acute to occasionally rounded, awned or long-mucronate, margins strongly and broadly recurved in proximal 4/5 of leaf, not bordered; costa long-excurrent, with an elongate adaxial pad of bulging, rounded cells with 3–4 simple papillae, distally narrow, 2–4 cells across adaxial surface; distal laminal cells subquadrate, 12–16 µm wide, 1: 1, smooth or less commonly papillose with 1 (–2) scattered simple papillae. Sexual condition autoicous. Sporophytes exerted. Seta ca. 0.4 (–0.9) cm. Capsule stegocarpic, not systylius, elliptic or cylindric, or erect and nearly straight, urn ca. 0.8–1.2 mm; peristome 100–150 µm, teeth 16, ligulate straight, or short, rudimentary, basal membrane lacking or very low; operculum 0.5 mm. Spores 15–18 µm, spheric, finely papillose.


Phenology: Capsules mature spring.
Habitat: Soil or walls, often in calcareous regions
Elevation: low to moderate elevations

Distribution

V27 845-distribution-map.gif

Ariz., Idaho, S.C., Tex., Europe, Asia, n Africa

Discussion

The tiny but often fruiting Tortula lanceola is uncommon in the flora area. The bulging, rounded cells of the adaxial costal pad have the distinctive appearance of cows’ udders, and viewed laterally have the appearance of longitudinal ridges. H. A. Crum and L. E. Anderson (1981) pointed out that the antheridia are borne naked in axillary clusters, but occasionally bracts, albeit rudimentary, are present. Tortula acaulon has much the same gametophyte, but the sporophyte is reduced and cleistocarpic. Tortula atrovirens and Crossidium aberrans have a somewhat similar leaf section but the former lacks an awn, and the latter has a short awn, and the adaxial costal pad is of low 1–2-celled filaments.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property."um" is not declared as a valid unit of measurement for this property.

... more about "Tortula lanceola"
smooth +  and papillose +
smooth +  and papillose +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
channeled +, nearly flat +  and concave +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
Richard H. Zander +  and Patricia M. Eckel +
R. H. Zander +
semicircular +  and elliptical +
semicircular +, elliptical +  and rounded +
papillose +  and smooth +
usually slightly wider +
usually ovate;oblong +
differentiated +
Encalypta lanceolata +
obovoid +
cucullate +
0.25 cm2.5 mm <br />0.0025 m <br /> (?) +  and 0.6 cm6 mm <br />0.006 m <br /> (?) +
not systylius +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
dark-brown +  and yellowish-brown +
cleistocarpous +  and stegocarpous +
cylindric;elliptic +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
monoicous +, dioicous +  and sexual +
twisted +, vesiculose +  and thicker-walled +
1 (?) +  and 3 (?) +
rounded +
tomentose +  and bare +
long-excurrent +
subpercurrent +
autoicous +  and sexual +
Ariz. +, Idaho +, S.C. +, Tex. +, Europe +, Asia +  and n Africa +
low to moderate elevations +
differentiated +
multicellular +
Soil or walls, often in calcareous regions +
somewhat larger +
yellow +  and orange-red or red +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
ovate +  and obovate +
reduced +
serrulate +  and entire +
smaller +
rostrate;long-conic +
0.5 cm5 mm <br />0.005 m <br /> (?) +
differentiated +
photosynthetic +
1 +, 3 +  and 4 +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
Capsules mature spring. +
Bull. Buffalo Soc. Nat. Sci. +
straight +
not twisted +  and twisted +
0.4 cm4 mm <br />0.004 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
brown;yellowish-brown +
elongate +
0.4 cm4 mm <br />0.004 m <br /> (?) +
2 +  and 1 +
orange +, red +, yellow +, laminal +  and light-brown +
spheric +
rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Pottia lanceolata +
Tortula lanceola +
species +
ovoid;cylindric +
straight +
rudimentary +
triangular +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
yellow distally and red +, orange +, red +  and yellow +
rectangular +
thickened +, thin +  and thicker +
stemless +  and perigoniate +
yellowbrown;dark-brown +
occasionally smaller than the perichaetiate +