Tortula leucostoma

(R. Brown) Hooker & Greville

Edinburgh J. Sci. 1: 294. 1824,.

Basionyms: Barbula leucostoma R. Brown Chlor. Melvill., 40. 1823
Synonyms: Desmatodon leucostoma (R. Brown) Berggren Desmatodon suberectus (Hooker) Limpricht Tortula suberecta Hooker
Treatment appears in FNA Volume 27. Treatment on page 599. Mentioned on page 589, 590.

Leaves ovatelanceolate or occasionally ovate, apex broadly acute or rounded-acute, piliferous, margins recurved in middle 2/3 of leaf or more, not or weakly bordered distally with ca. 2 rows of less papillose, thicker-walled cells, to ca. 2: 1; costa long-excurrent, lacking an adaxial pad of cells, distally narrow, 2–3 cells across the convex adaxial surface; distal laminal cells hexagonal, 13–20 µm wide, 1: 1, densely papillose with several 2-fid papillae. Sexual condition autoicous or paroicous. Sporophytes exerted. Seta 0.4–1.9 cm. Capsule stegocarpic, short-cylindric, erect and nearly straight, urn 1.5–2 mm; peristome 300–500 length µm, teeth 16, divided to near the base or perforate basally, the teeth often basally fused into a low basal membrane ca. 50 µm; operculum 0.5–0.6 mm. Spores ca. 23–25 µm, spheric, densely papillose.


Phenology: Capsules mature summer.
Habitat: Soil, silt, clay, especially calcareous substrates, occasionally runways and burrows of small mammals, typically subarctic, Arctic and alpine habitats
Elevation: low to high elevations (0-3700 m)

Distribution

V27 858-distribution-map.gif

Greenland, B.C., Nfld. and Labr., N.W.T., Nunavut, Yukon, Alaska, Ariz., Calif., Colo., Mont., Wash., Europe, Asia

Discussion

Tortula leucostoma and T. guepinii were considered by W. C. Steere (1939c) to be probably subspecies of T. hoppeana (as Desmatodon latifolius). This species differs from T. hoppeana largely by the tapering leaf with piliferous costa, and perhaps the somewhat shorter seta.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property.

... more about "Tortula leucostoma"
smooth +  and papillose +
smooth +  and papillose +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
Richard H. Zander +  and Patricia M. Eckel +
(R. Brown) Hooker & Greville +
semicircular +  and elliptical +
semicircular +, elliptical +  and rounded +
papillose +  and smooth +
usually slightly wider +
usually ovate;oblong +
differentiated +
Barbula leucostoma +
obovoid +
cucullate +
0.25 cm2.5 mm <br />0.0025 m <br /> (?) +  and 0.6 cm6 mm <br />0.006 m <br /> (?) +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
dark-brown +  and yellowish-brown +
cleistocarpous +  and stegocarpous +
short-cylindric +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
monoicous +, dioicous +  and sexual +
thicker-walled +
1 (?) +  and 3 (?) +
papillose +
elongate-rhomboid +  and elongate +
tomentose +  and bare +
long-excurrent +
subpercurrent +
paroicous +, autoicous +  and sexual +
Greenland +, B.C. +, Nfld. and Labr. +, N.W.T. +, Nunavut +, Yukon +, Alaska +, Ariz. +, Calif. +, Colo. +, Mont. +, Wash. +, Europe +  and Asia +
low to high elevations (0-3700 m) +
differentiated +
multicellular +
Soil, silt, clay, especially calcareous substrates, occasionally runways and burrows of small mammals, typically subarctic, Arctic and alpine habitats +
somewhat larger +
yellow +  and orange-red or red +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
ovate +  and ovatelanceolate +
reduced +
serrulate +  and entire +
smaller +
rostrate;long-conic +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
differentiated +
photosynthetic +
simple +  and hollow +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
Capsules mature summer. +
Edinburgh J. Sci. +
straight +
not twisted +  and twisted +
brown;yellowish-brown +
elongate +
0.4 cm4 mm <br />0.004 m <br /> (1.9 cm19 mm <br />0.019 m <br />) +
2 +  and 1 +
orange +, red +, yellow +, laminal +  and light-brown +
spheric +
rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Desmatodon leucostoma +, Desmatodon suberectus +  and Tortula suberecta +
Tortula leucostoma +
species +
ovoid;cylindric +
rudimentary +
divided +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
yellow distally and red +, orange +, red +  and yellow +
rectangular +
thickened +, thin +  and thicker +
stemless +  and perigoniate +
yellowbrown;dark-brown +
occasionally smaller than the perichaetiate +