Tortula nevadensis

(Cardot & Thériot) R. H. Zander

Bull. Buffalo Soc. Nat. Sci. 32: 226. 1993,.

Basionyms: Pottia nevadensis Cardot & Thériot Bot. Gaz. 37: 365, plate 18, fig. 1. 1904
Treatment appears in FNA Volume 27. Treatment on page 594. Mentioned on page 588, 589.

Leaves ovate to obovate or elliptical, apex acute, short-awned, sometimes long, margins plane or rarely weakly recurved medially, not bordered; costa long-excurrent, lacking an adaxial pad of cells, distally narrow, 2–3 cells across adaxial surface; distal laminal cells subquadrate or occasionally hexagonal, 16–22 µm wide, 1 (–2):1, smooth. Sexual condition autoicous. Sporophytes exerted. Seta 1–1.4 cm. Capsule stegocarpic, not systylius (but columella falls with operculum), obovate to cylindric, erect and nearly straight, urn 0.8–1.6 mm; peristome absent; operculum 0.6–1 mm. Spores 23–30 µm, spheric, granulose-papillose papillose.


Phenology: Capsules mature spring–summer.
Habitat: Soil, occasional saline soil, clay
Elevation: moderate to high elevations (500-3900 m)

Distribution

V27 848-distribution-map.gif

Alta., B.C., Ont., Que., Sask., Colo., Nev., Oreg., Utah, Wash.

Discussion

In Tortula nevadensis, the often long-cylindric capsule with operculum rostrate from a flattened base is much like that of Hennediella heimii but is not technically systylius because the operculum usually falls without being lifted by the columella; the columella is cut near the base and remains attached to the operculum. The leaves are reddish only in the extreme basal portion in 2 percent KOH solution. The costa seen in section has two (sub)stereid bands in the more robust leaves, but the large, smooth laminal cells and broad leaf shape preclude confusion with taxa in genera characteristically having two stereid bands (but only one band if plant environmentally depauperate). What appears to be a ventral stereid band may be secondary thickening of an internal epidermal layer. The distal leaves may be weakly denticulate near the apex by projecting cell ends. This uncommon species is a western North American one that is disjunctive to Ontario and Quebec.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property.

... more about "Tortula nevadensis"
smooth +  and papillose +
smooth +  and papillose +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
channeled +, nearly flat +  and concave +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
Richard H. Zander +  and Patricia M. Eckel +
(Cardot & Thériot) R. H. Zander +
semicircular +  and elliptical +
semicircular +, elliptical +  and rounded +
papillose +  and smooth +
usually slightly wider +
usually ovate;oblong +
differentiated +
Pottia nevadensis +
obovoid +
cucullate +
0.25 cm2.5 mm <br />0.0025 m <br /> (?) +  and 0.6 cm6 mm <br />0.006 m <br /> (?) +
not systylius +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
dark-brown +  and yellowish-brown +
cleistocarpous +  and stegocarpous +
obovate;cylindric +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
monoicous +, dioicous +  and sexual +
twisted +, vesiculose +  and thicker-walled +
1 (?) +  and 3 (?) +
elongate-rhomboid +  and elongate +
tomentose +  and bare +
long-excurrent +
subpercurrent +
autoicous +  and sexual +
Alta. +, B.C. +, Ont. +, Que. +, Sask. +, Colo. +, Nev. +, Oreg. +, Utah +  and Wash. +
moderate to high elevations (500-3900 m) +
differentiated +
multicellular +
Soil, occasional saline soil, clay +
somewhat larger +
yellow +  and orange-red or red +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
ovate +  and obovate or elliptical +
reduced +
serrulate +  and entire +
smaller +
rostrate;long-conic +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
differentiated +
photosynthetic +
simple +  and hollow +
4 +  and 6 +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
Capsules mature spring–summer. +
Bull. Buffalo Soc. Nat. Sci. +
straight +
not twisted +  and twisted +
brown;yellowish-brown +
elongate +
1 cm10 mm <br />0.01 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
2 +  and 1 +
orange +, red +, yellow +, laminal +  and light-brown +
spheric +
rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Desmatodon +, Phascum +, Pottia +  and Protobryum +
Tortula nevadensis +
species +
ovoid;cylindric +
rudimentary +
32 +  and 16 +
triangular +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
yellow distally and red +, orange +, red +  and yellow +
rectangular +
thickened +, thin +  and thicker +
stemless +  and perigoniate +
yellowbrown;dark-brown +
occasionally smaller than the perichaetiate +