Tortula obtusifolia

(Schwägrichen) Mathieu

Fl. Gén. Belgique 2: 48. 1853,.

Basionyms: Barbula obtusifolia Schwägrichen Sp. Musc. Frond. Suppl 1(1): 129, plate 31. 1811
Synonyms: Desmatodon arenaceus Sullivant Desmatodon coloradensis Grout Desmatodon obtusifolius (Schwägrichen) Schimper
Treatment appears in FNA Volume 27. Treatment on page 597. Mentioned on page 525, 588, 589, 590, 596, 599, 602, 636.

Leaves ovate to ligulate, apex rounded or occasionally broadly acute, apiculate, margins strongly recurved or revolute in distal 4/5 of leaf, sometimes bordered distally with (2–) 4 rows of thicker-walled cells, or occasionally border of rhomboidal, smooth cells commonly hidden in recurvature; costa ending in apiculus or occasionally percurrent or subpercurrent, lacking an adaxial pad of swollen cells or this poorly developed, distally not broadening, to 5–6 cells across the convex adaxial surface; distal laminal cells hexagonal or subquadrate, 10–13 µm wide, 1: 1, strongly papillose with 2-fid or multiplex papillae. Sexual condition autoicous, but plants often bearing only antheridia or archegonia. Sporophytes exerted. Seta 0.5–0.8 cm. Capsule stegocarpic, cylindric or ovate, erect and nearly straight, urn 1–2.5 mm; peristome rudimentary or to 500 µm, teeth 16, irregular, divided to base and filamentous when long, straight or twisted, basal membrane very low or absent; operculum 0.7–1 mm. Spores 9–11 (–15) µm, spheric, finely papillose or essentially smooth.


Phenology: Capsules mature spring–fall.
Habitat: Soil, rock, limestone, calcareous sandstone, stone walls, crevices, ledges
Elevation: low to high elevations (0-3300 m)

Distribution

V27 854-distribution-map.gif

Alta., B.C., Man., N.B., Ont., Que., Alaska, Ariz., Calif., Colo., Ga., Ind., Iowa, Kans., Mich., Minn., Mo., Mont., Nebr., N.J., N.Mex., Ohio, Okla., Pa., S.Dak., Tenn., Tex., Utah, Wash., Wis., Wyo., Europe, Asia, Africa

Discussion

The variation in peristome development of Tortula obtusifolia is considerable, and matches intraspecifically the variation between the peristomes of species previously recognized in the segregate genus Desmatodon and in Tortula in the strict sense. This species may be confused with T. atrovirens but the latter has a massive pad of bulging cells on the costa.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property.

... more about "Tortula obtusifolia"
smooth +  and papillose +
smooth +  and papillose +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
Acute (?) +  and Rounded (?) +
Richard H. Zander +  and Patricia M. Eckel +
(Schwägrichen) Mathieu +
semicircular +  and elliptical +
semicircular +, elliptical +  and rounded +
papillose +  and smooth +
usually slightly wider +
usually ovate;oblong +
differentiated +
Barbula obtusifolia +
obovoid +
cucullate +
0.25 cm2.5 mm <br />0.0025 m <br /> (?) +  and 0.6 cm6 mm <br />0.006 m <br /> (?) +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
dark-brown +  and yellowish-brown +
cleistocarpous +  and stegocarpous +
ovate;cylindric +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
monoicous +, dioicous +  and sexual +
thicker-walled +
5 (?) +  and 6 (?) +
swollen +  and rhomboidal +
broadening +
tomentose +  and bare +
subpercurrent +
Alta. +, B.C. +, Man. +, N.B. +, Ont. +, Que. +, Alaska +, Ariz. +, Calif. +, Colo. +, Ga. +, Ind. +, Iowa +, Kans. +, Mich. +, Minn. +, Mo. +, Mont. +, Nebr. +, N.J. +, N.Mex. +, Ohio +, Okla. +, Pa. +, S.Dak. +, Tenn. +, Tex. +, Utah +, Wash. +, Wis. +, Wyo. +, Europe +, Asia +  and Africa +
low to high elevations (0-3300 m) +
differentiated +
multicellular +
Soil, rock, limestone, calcareous sandstone, stone walls, crevices, ledges +
somewhat larger +
yellow +  and orange-red or red +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
reduced +
serrulate +  and entire +
smaller +
rostrate;long-conic +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
differentiated +
photosynthetic +
simple +  and hollow +
4 +  and 6 +
lateral +  and terminal +
lateral +  and terminal +
0-500 um +  and rudimentary +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
Capsules mature spring–fall. +
Fl. Gén. Belgique +
straight +
not twisted +  and twisted +
brown;yellowish-brown +
elongate +
0.5 cm5 mm <br />0.005 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
2 +  and 1 +
orange +, red +, yellow +, laminal +  and light-brown +
spheric +
rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Desmatodon arenaceus +, Desmatodon coloradensis +  and Desmatodon obtusifolius +
Tortula obtusifolia +
species +
ovoid;cylindric +
straight +
rudimentary +
divided +
0.1 cm1 mm <br />0.001 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
yellow distally and red +, orange +, red +  and yellow +
rectangular +
thickened +, thin +  and thicker +
stemless +  and perigoniate +
yellowbrown;dark-brown +
occasionally smaller than the perichaetiate +