Tortula porteri

(James) Brotherus in H. G. A. Engler and K. Prantl

in H. G. A. Engler and K. Prantl, Nat. Pflanzenfam. 214[I,3]: 430. 1902,.

Basionyms: Desmatodon porteri James in C. F. Austin, Musci Appalach., 123. 1870
Synonyms: Desmatodon fisherae H. A. Crum
Treatment appears in FNA Volume 27. Treatment on page 596. Mentioned on page 588, 590.

Leaves broadly ovatelanceolate, apex broadly acute or occasionally rounded, not or weakly apiculate, margins plane, always distinctly bordered nearly throughout with 2–4 rows of narrower, thicker-walled, smooth or weakly papillose cells; costa percurrent or ending in an apiculus, lacking an adaxial pad of cells, distally of even thickness, 4 (–6) cells across adaxial surface; distal laminal cells hexagonal or sometimes subquadrate, width 11–15 µm, 1: 1; hollow-papillose medially. Sexual condition dioicous. Sporophytes exerted. Seta 0.5–0.8 (–1) cm. Capsule stegocarpic, not systylius, cylindric, erect and nearly straight, urn 1.1–1.5 (–1.8) mm; peristome 200–350 µm, weakly twisted, teeth of 32 rather short filaments variously anastomosing proximally, basal membrane 60–100 µm; operculum 0.4–0.8 mm. Spores 8–10 µm, spheric, finely papillose or nearly smooth.


Phenology: Capsules mature spring.
Habitat: Limestone or dolomitic rock
Elevation: low to moderate elevations

Distribution

V27 853-distribution-map.gif

Ont., Que., Ark., Ill., Ind., Kans., Ky., Mo., N.J., Ohio, Okla., Pa., Tenn., Vt., Va., W.Va., South America (Venezuela)

Discussion

A report of Tortula porteri by Ced. L. Porter (1937) from Wyoming is doubtful. This species is quite like Hennediella in leaf traits such as plane margins, flattened superficial laminal cell walls, and border of elongate, thick-walled cells, but is yellow in KOH solution and lacks marginal teeth. The two genera intergrade somewhat and a less arbitrary distinction than simple emphasis on KOH color reaction awaits more study. This species is distinguished from T. obtusifolia by the more acute leaves with plane margins.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property."um" is not declared as a valid unit of measurement for this property.

... more about "Tortula porteri"
smooth +  and papillose +
smooth +  and papillose +
4 +  and lacking +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
channeled +, nearly flat +  and concave +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
Rounded (?) +  and Acute (?) +
Richard H. Zander +  and Patricia M. Eckel +
(James) Brotherus in H. G. A. Engler and K. Prantl +
semicircular +  and elliptical +
semicircular +, elliptical +  and rounded +
papillose +  and smooth +
usually slightly wider +
usually ovate;oblong +
differentiated +
Desmatodon porteri +
obovoid +
cucullate +
0.25 cm2.5 mm <br />0.0025 m <br /> (?) +  and 0.6 cm6 mm <br />0.006 m <br /> (?) +
not systylius +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
dark-brown +  and yellowish-brown +
cleistocarpous +  and stegocarpous +
cylindric +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
monoicous +, dioicous +  and sexual +
thicker-walled +
1 (?) +  and 3 (?) +
papillose +  and smooth +
elongate-rhomboid +  and elongate +
narrower +
tomentose +  and bare +
ending in an apiculus +  and percurrent +
subpercurrent +
dioicous +  and sexual +
Ont. +, Que. +, Ark. +, Ill. +, Ind. +, Kans. +, Ky. +, Mo. +, N.J. +, Ohio +, Okla. +, Pa. +, Tenn. +, Vt. +, Va. +, W.Va. +  and South America (Venezuela) +
low to moderate elevations +
differentiated +
multicellular +
Limestone or dolomitic rock +
somewhat larger +
yellow +  and orange-red or red +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
ovatelanceolate +
reduced +
serrulate +  and entire +
smaller +
rostrate;long-conic +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
differentiated +
photosynthetic +
simple +  and hollow +
4 +  and 6 +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
Capsules mature spring. +
in H. G. A. Engler and K. Prantl, Nat. Pflanzenfam. +
straight +
not twisted +  and twisted +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
brown;yellowish-brown +
elongate +
0.5 cm5 mm <br />0.005 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
2 +  and 1 +
orange +, red +, yellow +, laminal +  and light-brown +
spheric +
rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Desmatodon fisherae +
Tortula porteri +
species +
ovoid;cylindric +
rudimentary +
32 +  and 16 +
triangular +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
yellow distally and red +, orange +, red +  and yellow +
rectangular +
thickened +, thin +  and thicker +
stemless +  and perigoniate +
yellowbrown;dark-brown +
occasionally smaller than the perichaetiate +